EFFECT OF FRESHWATER INFLOW ON MACROBENTHOS PRODUCTIVITY AND NITROGEN LOSSES IN TEXAS ESTUARIES Paul A. Montagna, Principal Investigator TWDB Contract No. 94-483-003 Technical Report Number TR/94-005 OCTOBER 1994 FINAL REPORT EFFECT OF FRESHWATER INFLOW ON MACROBENTHOS PRODUCTIVITY AND NITROGEN LOSSES IN TEXAS ESTUARIES by Paul A. Montagna, Principal Investigator from University of Texas at Austin Marine Science Institute P.O. Box 1267 Port Aransas, Texas 78373 to Texas Water Development Board 611 South Congress Avenue, Suite 116 Austin, Texas 78704-1736 Interagency Cooperative Contract TWDB Contract No. 94-483-003 The University of Texas Marine Science Institute Technical Report Number TR/94-005 TABLE OF CONTENTS LIST OF CONTRIBUTIONS ii Scientific Publications ii Technical Reports ii Invited Workshops ii Invited Seminars iii Contributed Oral Presentations at National Meetings iii PREFACE iv ACKNOWLEDGEMENTS v ECOLOGY OF INFAUNAL MOLLUSCA IN SOUTH TEXAS ESTUARIES 1 RECENT BIOLOGICAL DATA COLLECTIONS 47 HYDROGRAPHIC DATA 47 NUTRIENT CONCENTRATIONS 52 BIOMASS DATA 55 SPECIES DATA 76 NITROGEN LOSSES TO ESTUARINE SEDIMENTS 118 . LIST OF CONTRIBUTIONS All of the following scientific contributions have been performed during the contract reporting period (September 1, 1993-August 31, 1994) and acknowledged support, or partial support, by the Texas Water Development Board, Water Research authorized under the Texas Water Code sections 15.402 and Planning Fund, administered by the department under interagency 16.058(e). This support was cooperative contract numbers: (1986-87) 0757, 8-483-607, 9-483-705, 90-483-706, 91-483-787, 92-483-300, 93-483-352, and 94-483-003. Scientific Publications Koepfler, E.T., R. Benner, and P.A. Montagna. 1993. Variability of dissolved organic carbon in sediments of a seagrass bed and an unvegetated area within an estuary in southern Texas. Estuaries 16:391-404. Montagna, P.A., Stockwell, D.A., and Kalke, R.D. 1993. Dwarf surfclam Muiinia lateralis (Say, 1882) populations and feeding during the Texas brown tide event. Journal of Shellfish Research 12:433-442. Technical Reports Montagna, P.A. 1994. Inflow needs assessment: Effect of the Colorado River diversion on benthic communities. Final Report to the Lower Colorado River Authority. Technical Report No. TR/94-001, Marine Science Institute, The University of Texas, Port Aransas, TX 63 pp. Invited Workshops (*indicates invited presentation) *Minorities in Marine Science. September 11, 1993. University of Texas Marine Science Institute, Port Aransas, Texas. Lavaca Bay Scientific Workshop. February 3-5, 1994. Texas General Land Office. Flouston, Texas. *Gulf Intracoastal Waterway dredge disposal issue. March 7, 1994. Coastal Bend Bays Foundation. Corpus Christi, Texas. Invited Seminars "Crash of benthic communities." Port Montagna, P.A. Turning the tide symposium. Aransas, Texas. August 20, 1994. Montagna, P.A. and R.D. Kalke. "Ecology of infaunal Mollusca of south Texas estuaries". American Malacological Union. Houston, Texas. July 9-14, 1994. Montagna, P.A. "Benthic communities and dredging." Lower Laguna Madre Conference. November 20, 1994. Harlingen, Texas. Montagna, P.A. "Relationship between climate, freshwater inflow, and benthos in Texas estuaries." Estuarine Research Federation Conference. Hilton Head, South Carolina. November 14-19, 1993. Montagna, P.A. A.F. Amos, R. Benner, E.J. Buskey, K.H. Dunton, P.L. Parker, D. Stockwell, and T.E. Whitledge." An ecosystem study of Laguna Madre, Texas. Estuarine Research Federation Conference. Hilton Head, South Carolina. November 14-19, 1993. Montagna, P.A. "Life in the mud of Texas Bays." Dean’s Scholars Workshop. Port Aransas, Texas. October 30, 1993. Montagna, P.A. "Estuarine and Benthic Research." Minorities in Marine Science Workshop. Port Aransas, Texas. September 11, 1993. Contributed Ora! Presentations at National Meetings Martin, C. and Montagna, P.A. LaQuinta Channel environmental monitoring project; benthic diversity. South Texas Bays and Estuaries Meeting. Port Aransas, Texas. April 1, 1994. PREFACE The primary goal of the current research program is to define quantitative relationships between marine resource populations and freshwater inflows to the State’s bays and estuaries. However, we know that there is year-to-year variability in the population densities and successional events of estuarine communities. This year-to-year variability is apparently driven by long-term, and global-scale climatic events that affect the rates of freshwater inflow. Therefore, this report documents long-term changes in populations and communities that are influenced by freshwater inflow. The best indicator of productivity is the change in biomass of the community. A Secondary goal of the current research program is to quantify the loss of nitrogen in Texas estuaries. Nitrogen is the key element that limits productivity. A simple budget would account for nitrogen entering the bay via freshwater inflow, how it is captured and transformed into biomass, and finally how it is lost from the ecosystem. One aspect of nitrogen loss is very poorly understood: "How much nitrogen is buried and lost from the system?" We report here nitrogen content changes with respect to sediment depth. Presumably nitrogen is labile in the upper, biologically active, layers of sediment, and refractory at depth. Therefore, it is important to determine the sediment depth at which nitrogen content is at a low and constant value. This study is a continuation of freshwater inflow studies that began in 1984. The goals have evolved over the years to reflect the synthesis of new information and the management needs of the Texas Water Development Board. The original studies (1984-1986) were designed to determine the effect of inflow on Lavaca Bay. One station used during that study is still being sampled. San Antonio Bay was studied in 1987, and the Nueces Estuary (Nueces and Corpus Christi Bays) were studied in 1988. Long-term studies of the Lavaca-Colorado and Guadalupe Estuaries began in 1990. This research has been supplemented by other projects. For example, studies of the Laguna Madre were made possible by funded by the Texas a program Advanced Technology Program. The Lower Colorado River Authority recently supplemented the long-term study in Matagorda Bay by adding funding to sample additional stations in the Eastern arm of Matagorda Bay to study the effects of the diversion of the Colorado River. Long-term studies on the Nueces Estuary have been recently funded by the Texas Sea Grant Program. The new Sea Grant project is utilizing stations originally established by the Texas Water Development Board projects in 1988. The primary focus of the Sea Grant program is to determine the role of climatic variability in controlling productivity in estuaries. Although, there is ten years worth of data in some cases, we have not sampled over two entire wet-dry cycles. We have sampled over one and one-half cycles. We are currently beginning to enter a dry cycle. The completion of this research will take about two more years and should end when we enter the next wet cycle, which will be heralded by the next El Nino event. The structure of this report is as follows. Chapter one is a manuscript which This has been submitted for publication in the American Malacological Bulletin. manuscript summarizes all biological data on mollusks taken to date and synthesizes current information on the biological effects of freshwater inflow on a key indicator taxanomic group. Chapter two is a compilation of biological and hydrographical data obtained during the last two years. Chapter three is a compilation of all the sediment data on nitrogen losses that has been collected to date. Chapter three also contains brief narrative describing the data and drawing preliminary conclusions. a ACKNOWLEDGEMENTS I must acknowledge the significant contributions of Mr. Rick Kalke. Rick began 1984. and the first sampling study of Lavaca Bay in He is an outstanding field person taxonomist. The work reported on in this study could not have been performed without him. Carroll Simanek also provided significant help in data management. We obviously are collecting and processing a large amount of data. Input, proof-reading and maintenance of this large data set is a daunting task that Carroll handles very well. This work has also benefitted by discussions with colleagues at agencies. Gary Powell, William Longley, and David Brock of the Texas Water Development Board have provided much help and guidance. Recently, Cynthia Gorham of the Lower Colorado River Authority has also been supplying us with help on the Matagorda Bay study area. ECOLOGY OF INFAUNAL MOLLUSCA IN SOUTH TEXAS ESTUARIES by Paul A. Montagna and Richard D. Kalke Abstract: The ecology of Texas estuaries is strongly influenced by latitudinal ecotones that exist along the Northwestern Gulfof Mexico coastline. Long-term studies have been conducted in four of the seven major estuarine ecosystems in Texas. The objective is to determine the role of climatic variability and concordant differences in freshwater inflow among the ecosystems in structuring benthic infaunal communities and maintaining secondary production. Mollusks are prominent members of the infauna in all benthic habitats of Texas estuaries. The abundance, biomass and community structure of mollusks was measured along salinity gradients within the four south Texas estuaries. Overall, a Texas estuary has on average 14 species of mollusks, with an 22 average abundance of 7,500 individuals-m’, and an average biomass of 2.4 g-m' . Freshwater inflow is the dominant factor regulating variability of molluscan communities. Salinity is a surrogate for inflow, therefore, there are within and among estuarine zoogeographic patterns related to salinity patterns. There are seasonal, interannual and latitudinal patterns of inflow, and these patterns are apparently regulating community structure, population dynamics and secondary production in Texas estuaries. Recent water projects to enhance the amount of freshwater flowing into estuaries appear to have had an effect and have increased the number of mollusks in those areas. However the projects occurred during a naturally wet period, so it is difficult to differentiate natural versus anthropogenic changes. The response of mollusks to natural gradients and man- induced changes of freshwater inflow demonstrate the importance of this factor in regulating benthic communities. MONTAGNA AND KALKE; TEXAS INFAUNAL MOLLUSCA A major component of benthic ecosystems in Texas estuaries, as is true elsewhere, are the Mollusca. Molluscan biomass dominates the macroinfauna in Lavaca, San Antonio Corpus Christi, and Nueces Bays (Kalke and Montagna, 1991; Montagna and Kalke, 1992). During peak recruitment events, mollusca can also dominate population abundance. However, differences in population size and community structure exists within and among Texas Bays. There are seven major estuarine systems along 373 linear miles of coastline. The estuaries of Texas are remarkably diverse in spite of similar physiography (Fig. 1). This is due to a climatic gradient, which influences freshwater inflow. The gradient of and concomitant freshwater inflow, from north to south, is the most decreasing rainfall, distinctive feature of the coastline (Table 1). Along this gradient, rainfall decreases by a factor of two, but inflow balance decreases by almost two orders of magnitude. The inflow patterns appear to group into four distinct types of estuaries that vary by about an order of magnitude each (Table 1). Each estuary-type also has distinctly different timing of peak inflow events. The northern estuaries receive peak inflow during the spring, the central estuaries are bimodal receiving peak inflows during the spring and fall, and the southern most estuaries receive peak inflows during the fall (Texas Dept. Water Resources, 1982). These distinct patterns are very important, since growth, reproduction, and migration of many species is keyed to seasonal events. The timing and magnitude to regulate production of inundation is believed finfish and shellfish (Texas Dept. Water Resources, 1982). We have been conducting long-term studies in four of the seven estuaries to determine the role of freshwater inflow in maintaining benthic productivity. The primary purpose of the current study is to determine the degree of influence of freshwater inflow in regulating zoogeographic differences of molluscan population size and community structure within and among Texas estuaries. The secondary purpose of this study is to assess the effects of two major water projects designed to increase freshwater inflows to estuaries to maintain or enhance productivity. One project is a mandated freshwater MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA release schedule from a dam and the other is a diversion of river water to an estuary. The focus in this manuscript is on the infaunal mollusks. METHODS All seven Texas estuaries have similarities in their structure and physiography (Fig. 1). A barrier island runs parallel with the entire length of the coast. Between the island and the mainland there are lagoons. The lagoons are interrupted with drowned river valleys which form the bay and estuarine systems. There is a Gulf inlet through the barrier island, connecting the sea with the lagoon behind the island. The lagoon opens to a large primary bay. There is a constriction between the primary bay and the smaller secondary bay. The river flows into the secondary bay. Primary bays have greater marine influence and secondary bays have greater freshwater influence. So, as well as a latitudinal climatic gradient, there is a longitudinal salinity gradient within each estuary. The similarity of the Texas estuaries allows us to design a sampling program where we can use statistical control on confounding factors, e.g., Gulf exchange, circulation patterns, and alterations by man. Four to six stations were chosen in each I estuary (Table 2, Fig. 1), employing the same spatial sampling design have employed in previous studies of Texas estuaries (e.g., Montagna and Kalke, 1992)! Two replicate stations (A and B) are in the secondary bay where freshwater influences are greatest. Two other replicate stations (C and D) are in the primary bay where marine influences are greatest. By using two stations in the freshwater influenced zone and two stations in the marine influenced zone we are replicating effects at the treatment level and avoiding pseudoreplication (Hurlbert, 1984). There has been a diversion of the Colorado River into the east arm of Matagorda Bay, so we have located two additional station (E and F) there. The stations in Laguna Madre are located using a similar strategy. Two stations are located in Baffin Bay (6 and 24), and two station are located in Laguna Madre in a seagrass bed (189G) and an unvegetated sand patch (1898) (Fig. 1). Two major water projects were initiated during the course of this study. The MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA of both projects was to increase freshwater inflows to bays in order to enhance secondary productivity. In 1990, the Texas Water Commission ordered The City of Corpus Christi to release 151,000 ac-ft/y (1.86*10 8 m^y'1 ) to the Nueces Estuary from the Choke Canyon/Lake Corpus Christi reservoir system. The releases were mandated, because The City had not been releasing water. Stations A and B in Nueces Bay are purpose used to assess the effects of this project (Fig. 1). The Colorado River was diverted into the eastern arm of Matagorda Bay by the creation of a flood diversion channel in 1991 and a dam in the river channel below the point of diversion in 1992. This project has diverted Colorado River water from the Gulf of Mexico into the eastern arm of Matagorda Stations E and F were sampled to assess the effect of this diversion into Bay. Matagorda Bay (Fig. 1). The current study is not a complete assessment of the efficacy of these two projects. Three replicate sediment samples were taken within a 2 m radius at each of the stations in each estuary four times per year. Abundance and community structure were measured using the standard techniques that we (Montagna and Kalke, 1992) have been using since 1984. This includes sectioning 6.715 cm diameter cores (at 0-3 cm, and 3­10 cm) to examine the vertical distribution of infauna. Animals are then extracted using a 0.5 mm sieve, enumerated, and identified. The taxonomic authorities were Abbott (1974) and Andrews (1992). Principal components analysis was performed on all data sets to determine the relationship among stations in terms of species composition. Hydrographic data was recorded at each station using a Hydrolab Surveyor 11. These measurements include: salinity, conductivity, temperature, dissolved oxygen, ORP, pH, and depth. RESULTS SALINITY REGIMES MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 2-5). 1985-1986 and 1992-1993 were wet periods with concordant low salinities. These wet periods occur during periods when an El Nino is occurring in the western Pacific Ocean. The intervening time between El Nino events is dry. Texas suffers through a series of flood and drought periods, which is being regulated by global climatic events. Within each there are generally lower salinities in the spring Seasonality exists. year, and higher in the summer. Salinity in the Lavaca-Colorado Estuary ranges from 0-36 psu (Fig. 2). The lowest salinities always occur in the secondary bay at stations A and B. After the diversion of the Colorado River, Stations E and F exhibited low salinities that are more typical of the secondary bay. Salinity in the Guadalupe Estuary ranges from 0-32 psu (Fig. 3). During flood periods, this estuary is uniformly low (0-10 psu) in salinity. This is unusual compared to other Texas estuaries. It is caused by the high rate of inflow into a relatively small estuary (Table 1). The high turnover rate and low rate of exchange of marine water with the Gulf of Mexico exacerbate this trend. During extreme flooding the entire estuary can be at or near 0 psu. During drought periods, there can be a gradient of salinity. Salinity in the Nueces Estuary ranges from 2-45 psu (Fig. 4). Prior to 1991, salinities in the estuary were uniform and high. In 1991, a series of mandated freshwater releases began. This resulted in lowered salinities in the secondary bay. Salinities in the secondary bay were much lower than in the primary bay, where they had be similiar in 1987-1988. Heavy rain in 1992-1993 reduced salinity further. from Seasonal Salinity in the Laguna-Baffin Estuary range 10-60 psu (Fig. 5). fluctuations is less evident in the system. Changes occur system-wide when there are large climatic events, e.g., the 1992-1993 El Nino. There is little salinity gradient in this ecosystem, because freshwater inflow and exchange with the Gulf of Mexico is restricted. COMMUNITY STRUCTURE In the Lavaca-Colorado Estuary, stations A and B are almost identical (Fig. 6). MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Station F, at the mouth of the river diversion is also similar to A and B. Stations C and E are similar, and both these stations are nearly equi-distant from freshwater input and Gulf exchange. Station D, near the pass, is the most different station of all. The pattern elucidated in the principal components analysis is driven by the greater number of species that were found in station D, near the Gulf pass (Table 3). Also, species dominance patterns are different. The dominant pelecypods are from the genus Periploma at stations C and D, whereas Mu/in/a lateralis is the dominant at stations A, B, E and F where freshwater influence. Gastropod species are more uniformly distributed throughout the estuary. The dominant gastropods were Nassarius acutus and Acteocina candei. Pelecypods were always dominant over gastropods. Gastropods were most common in Lavaca Bay where they constituted 32% of the population at station A and 49% at B. In contrast, gastropods represented only 24% in C, 14% in D, 16% inEand 20%in F. In the Guadalupe Estuary, all of the stations are somewhat alike in terms of community structure (Fig. 7). There is more of a gradient from stations A to B to C to D in terms of abundance of individual species (Table 4). This is true for the dominant species, e.g., Texadina sphinctostoma, Acteocina candei and Muiinia lateralis. The f brackish species, Rangia cuneata only occurs in stations A and B. In general, there are more species in the marine end of the estuary where stations C and D are located. However, there are much higher abundances of species in the freshwater end of the estuary where stations A and B are located (Table 4). In spite of these trends, The principal components analysis indicates that there may be more affinity between stations A and C, and stations B and D may be more alike (Fig. 7). This trend may be explained by the unusual circulation pattern in San Antonio Bay. Freshwater enters the estuary near station A, and travels southwest along the shoreline toward station C. Marine water enters the Bay near station D and travels north toward station B. The species community pattern in the principal components analysis is driven by the number of Gastropods the number of pelecypods. Gastropods most at versus were common station A (80% of the population) and station C (58%). In contrast, gastropods MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA In the Nueces Estuary, stations A and B in Nueces Bay are almost identical (Fig. 8). Stations D and E are very similar, and station C is somewhat different from all other stations. Stations D and E are nearest the Aransas Pass in Corpus Christi Bay. Station C is in the upper part of Corpus Christi Bay. The pattern elucidated in the principal components analysis is driven by the greater number of species that were found in stations D and E, near the Gulf pass (Table 5), and some species unique to station C. In stations A and B, the dominant species are the pelecypods, Muiinia lateralis and Macoma mitchelli. In contrast, at stations D and E, gastropods were always dominant, where they constituted 46% of the population at station D and 33% at E. Gastropods represented only 7% in A and B, and 16% in C. Station C was different from the rest in that the dominant pelecypods were from the family Nuculanidae (Table 5). The Laguna Madre-Baffin Bay system exhibited the most varied molluscan communities within an ecosystem (Fig. 9). This trend was due to the difference caused habitats of Laguna Madre, versus the open Bay habitats characteristicby the seagrass of Baffin Bay. Stations 189 G and 189 S were identical, and stations 6 and 24 were identical (Fig. 9). Gastropods are rich in Laguna Madre (13 species in stations 189G and 11 species in 189S), but few in Baffin Bay (3 species at station 6 and 24) (Table 6). Only one pelecypod species, Muiinia lateralis was ever found in Baffin Bay. In contrast, , 11 species were found in station 189 G and 9 species were found in station 189S. Because of the concomitant low numbers of individuals found in Baffin Bay, the proportion of each class was similar. Gastropods dominate this ecosystem, 77% at station 189G, 71% at 1898, 67% at 6 and 38% at 24. Community characteristics vary among the estuaries as well as within the estuaries (Table 7). Salinity generally increases from north to south. The lowest salinity, open bay station (A in the Guadalupe) has the highest abundance and biomass indicating high productivity. The only high salinity station with high abundance and biomass is the seagrass habitat of Laguna Madre (station 189G). The hypersaline environments of Baffin Bay have the lowest abundances and biomasses. Diversity is generally highest near Gulf passes and in the seagrass habitats. Overall, a Texas estuary has on average 14 species of mollusks, with an average abundance of 7,500 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 22 individuals-m'and 2.4 g*m' . , The molluscan community in Texas estuaries is dominated by the dwarf surf clam, Mulinia lateralis (Table 8). Mulinia lateralis populations are more abundant in the fresher bays of the northern part of the study area. Only in Laguna Madre, which is dominated by seagrass beds, were other species found to dominate. Salinity alone is not the only determining factor. Estuarine physiography is also very important. For example, Mulinia lateralis is found in Lavaca Bay and Matagorda Bay, but is at highest density in stations C and E, which have moderate inflow influence, and moderate average salinities near 23 psu (Table 3). But, in the higher salinity Baffin Bay (about 41 psu), it is still the dominant species (Table 6). Mulinia lateralis is abundant in the Guadalupe Estuary where salinities from 7-16 but is most abundant in range psu, stations A and B where the saliinity is low, about 9 psu (Table 4). Mulinia lateralis is also equally present in most stations in the Nueces Estuary, where average salinities range from 23 to 32 psu (Table 5). On average, Mulinia lateralis is more dense in the The secondary bays than in the primary bays from San Antonio Bay south to Baffin Bay. only exception is in the Matagorda Bay, but here it also occurs nearer the freshwater inflow sources, and is rare near the Gulf pass. There is a great deal of temporal variability with respect to the densities of all these organisms. However, it is most exemplified by Mulinia lateralis (Fig. 10). The temporal patterns within estuaries are similar, but are of different magnitude. 1987-1988 and 1992-1993 Mulinia lateralis in the central estuaries (Fig. 10A). were good years for This pattern was not as distinct in the southern estuaries (Fig. 10B). The good years correspond to wet years which followed, or occurred during El Nino events. Mulinia lateralis practically disappeared from Baffin Bay during 1990-1992 corresponding to the occurrence of a severe brown tide bloom. Large-scale climatic events seem to be controlling Mulinia lateralis populations. MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA DISCUSSION On one hand, there appears to be a typical open bay "Texas molluscan" community. The community is dominated by small bivalves. Typically the small bivalves represent two-thirds of the community. The dominant species are Mulinia lateralis and Macoma mitchelli. The estuarine-wide pattern is influenced by the patterns the near river mouth, where bivalves can be as high as 90% of the population. The dominance of these clams is important to the entire trophic structure of Texas estuaries, because M. lateralis is the predominant food source for black drum, Pogonias cromis (Martin, 1979). There are two exceptions to this "generalized Texas community," where gastropods are dominant: San Antonio Bay and Laguna Madre. Laguna Madre is dominated by seagrass bed habitats, and diversity and standing stocks are generally very high. This is a direct result of the value of the seagrass habitats. San Antonio Bay is unusuall in that it is dominated by freshwater inflow. This is due to high turnover rates of water and low rates of marine exchange. It appears that hydrography and physiography are responsible for the different kind of community found in San Antonio Bay. During the entire current study period, except for 1989, there has been a continuous brown tide bloom in Laguna Madre and Baffin Bay (Stockwell et aL, 1993). on Other brown tide blooms have had catastrophic effects bivalves (Shumway, 1990). Effects have ranged from reproductive or recruitment failures (Bricelj et aL 1987; Tracey, , 1988), to adverse effects on feeding (Bricelj and Kuenstner, 1989; Tracey, 1988, Tracey etaL toa etaL 1988) toxic effect (Draper 1989; Tracey et aL, 1990; Gainey and Shumway, 1991). So, it is possible that the trend reported here is not normal, and may be due to the effects of a brown tide. We know that M. lateralis feeds on brown tide ,, (Montagna et a/. 1993), so it is likely that the changes in Baffin Bay are related to an f effect on larvae, reproduction, completely unrelated to brown tide. If the brown or are tide has had an effect on bivalves in Laguna Madre, then the conclusion that gastropods MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA brown tide. Brown tide is not a possible explanation of the dominance by gastropods in San Antonio Bay. In fact, San Antonio Bay has a large population of M. lateralis (Fig. 10). It is just that the population of Texadina sphinctostoma is enormous at the mouth of the Guadalupe River (Table 4). The undentified gastropods in these samples are all very small juveniles, and are also likely to be Texadina sphinctostoma. San Antonio Bay is also less like other Texas bays in terms of physiography. The demarcation between the primary and secondary bay is less distinct (Fig. 1), and there is very indirect exchange between the Bay and the Gulf of Mexico. In San Antonio Bay most living freshwater found in the absent from the eastern shore (Parker, 1959). The distribution of Rangia cuneata in species are upper bay and along the west shoreline being conspicuously upper San Antonio Bay and along the west shoreline conforms with the dominant freshwater flow pattern (Ladd, 1951). In general, the average pattern in the Guadalupe Estuary is masked by an unusual pattern of freshwater species near the River and the circulation pattern. the have demonstrated the importance of salinity Ecological studies over years factor in affecting the distribution of marine and estuarine organisms. The number as a of species, but not necessarily the observed total biomass increases as one proceeds along a salinity gradient from the freshwater side of a large estuary to the open sea (Springer and Woodburn, 1960; Gunter, 1961). This trend is also evident among Texas Mollusca (Table 7). The dominant species found in the current study are Mulinia lateralis and Texadina sphinctostoma (Table 8). The distribution of these species is strongly linked to long term environmental conditions, although responses to flood conditions may result in rapid population changes. Texadina sphinctostoma , a gastropod, populations increase following peaks in freshwater inflow (Harper, 1973; Matthews et a!., 1974). This is apparently a breeding response caused by a salinity decline (Harper, 1973). Texadina carries its eggs on the shell and undergoes direct development with the young ready to assume adult existence upon emerging from the egg. Texadina sphinctostoma is commonly reported as one of the most dominant gastropod inhabitants of the river influenced upper bays of the Texas MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA coast (Ladd, 1951; Ladd et al., 1957; Parker, 1959; Harper, 1973; Matthews et a/., 1974; Gilmore et a/., 1976; White et al., 1983; 1989; Staffers/., 1985). Mulinia lateralis is an extremely hardy species, ranging from Prince Edward Island, Canada to Yucatan, Mexico and in salinities from 5 ppt to 80 ppt (Parker, 1975). It is an opportunist of adversity because it can colonize rapidly after a disturbance event such as dredging or heavy rain (Flint and Younk, 1983; Flint et a!., 1981). It is one of the more abundant mollusks in the low salinity bay heads of the Gulf coast (Hopkins et a!., 1973). In San Antonio Bay Matthews et al. (1974) and Harper (1973) reported Mulinia widely distributed from the brackish water to higher salinity as found here. Both authors indicated that the close resemblance of Rangia juveniles and Mulinia lateralis have resulted in numerous misidentifications at the low salinity stations. In the may Laguna Madre (Alazan Bay) Mulinia lateralis was the most abundant and widespread mollusc (Martin 1979, Cornelius 1984). Mulinia lateralis is widely reported from other bays around the globe. Spawning was observed in the Tred Avon River, Maryland and Chesapeake Bay where it was observed to have a continuous period of setting from a single spawning cycle from May through November (Shaw, 1965; Holland et a!., 1977). In Alazan Texas Cornelius (1984) observed juveniles in all months Bay, except December, and Poff (1973) observed year round spawning in Trinity Bay, Texas. Mulinia lateralis has a very short generation time and is capable of successfully spawning at 3 mm in length which is approximately 60 days old (Calabrese, 1969a). Embryo survival and development for Mulinia as it is with Rangia cuneata is dependent on certain salinity and temperature ranges. Mulinia lateralis developed into normal larvae throughout the salinity range of 15 to 35 ppt and the temperature range of 10 to 30°C (Calabrese, 1969b). This clam is an important food item to bottom feeding organisms, i.e., the black drum (Pearson, 1929; Breuer, 1957; Simmons and Breuer, 1962; Martin, 1979) and to the greater and lesser scaup ducks (Cronan, 1957). Large rafts of ducks were observed in San Antonio in November 1988 scaup upper Bay corresponding to high densities of Mulinia lateralis (personal observation). MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA is no exception. Recently, intervention has been attempted by State agencies to try and conserve or enhance natural resources. Two projects occurred during the current study; a mandated freshwater release schedule in Nueces Bay, and a diversion of the Colorado a River to the east arm of Matagorda Bay. Both projects appear to have had desirable effect, if this is defined as enhanced molluscan communities. The physical effect of both projects is obvious from the lowered salinity values that have occurred (Figs. 2 and 4). In the Nueces Estuary, the stations A and B, had very high salinities, and no longer exhibit this characteristic. There were generally, low abundances and diversity prior to the mandated releases, and there is now a productive community. In the Lavaca-Colorado Estuary station F is now very much like Station A in terms of salinity and species composition. Both projects appear to have had the desired effects. However, there are two caveats. The changes that have occurred in the Nueces Estuary happened during an El Nino event, and therefore may have occurred without the releases. The changes that have occurred in the Lavaca-Colorado Estuary are also being mitigate by siltation, which threatens to close the diversion or restrict freshwater inflow. Only after several more years of sampling, to determine the long-term effect, can we be certain that these two projects will have lasting value. Freshwater inflow has obvious benefits to estuaries. Only Texas estuaries with high freshwater inflow rates support a productive shellfish industry (Table 1). The distribution and abundance of M. lateralis demonstrates the importance of inflow and the physical characteristics of estuaries. The variability of salinity patterns is more important than the absolute salinity values. Mulinia lateralis occurs most frequently in Baffin Bay, which is hypersaline, and Lavaca Bay and San Antonio Bay, which both low salinity are regions. However, all three bays are secondary bays. This means that freshets have large impacts on these systems, and salinity values change rapidly. If only absolute values of salinity were important, then these patterns would not exist. It is more likely that recruitment events for M. lateralis are initiated by a large change in the salinity occur value, which predominantly in secondary bays. Mulinia lateralis is apparently a good indicator species of freshwater inflow effects. MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA freshwater inflow, estuarine physiography, and time. Obviously, biological interactions are also occurring. However, we have little information to determine how to separate effects due to physical factors and biological interactions, e.g., competition and predation. There are obviously interactions among three geophysical factors, which regulate and control the structure and function of molluscan communities in Texas estuaries. These factors are: climate (which regulates rates of freshwater inflow), estuarine physiography (which regulates circulation patterns and the degree of marine exchange), and the presence ofspecific habitats (particularly seagrass beds). These factors control the landscape of the estuary and this determines both the makeup and productivity of the molluscan community. Climate and physiography interact to control the salinity patterns among and within the estuaries. The salinity patterns are good surrogates for indicating the effects of climate and physiography, but salinity itself is not the controlling factor. It is clear that freshwater inflow is very important in maintaining estuarine productivity. The potential for enhancing marine resources by water management projects appears to be a fruitful endeavor, but this must be confirmed with long-term ecosystem level research. ACKNOWLEDGEMENTS This manuscript was the result of many different research projects. The research in the Lavaca-Colorado, Guadalupe and Nueces Estuaries was partially funded through the Texas Water Development Board’s (TWDB) Water Research and Planning Fund, authorized under Texas Water code Sections 15.402 and 16.058(e), and administered by the Department under interagency cooperative contracts Nos. 8-483-607, 9-483-705, 9-483-706, 93-483-352 and 94-483-003. The authors have benefitted by discussions with Gary Powell, William Longley, and David Brock of the TWDB. Other partial support for work in Matagorda Bay was supplied by the Lower Colorado River Authority (LCRA). The authors have benefitted by discussions with MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Cynthia Gorham of the LCRA. The work in the Nueces Estuary was partially funded by Institutional Grant NA16RG0445-01 to Texas A&M University Sea Grant Program from the National Sea Grant Office, National Oceanic and Atmospheric Administration, U.S. Department of Commerce. Partial support was provided forwork in Laguna Madre and Baffin Bay by the Texas Higher Education Coordinating Board, Advanced Technology Program under Grant No. 4541 (and 3658-264). Partial support was also provided by University of Texas at Austin, Marine Science Institute. The author especially thanks Ms. Carol Simanek for playing a vital role in data management. Anotonio Mannino, Chris Martin, Rob Rewolinsky, Amy Rutter and Greg Street helped with sample analysis. 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The Influence of Freshwater Inflows Upon the Major Bays and Estuaries of the Texas Gulf Coast. Vol. 8. Executive Summary (Second Ed.). Texas Department of Water Resources, Austin, Texas. 133 pp. Texas Parks and Wildlife. 1988. Trends in Texas Commercial Fishery Landings , 1977-1987. Management Data Series, No. 149. Texas Parks and Wildlife Texas. Department, Coastal Fisheries Branch. Austin, Tracey, G.A. 1988. Feeding reduction, reproductive failure, and mortality in Mytilus edulis during the 1985 ’brown tide’ in Narragansett Bay, Rhode Island. Mar. Ecol. Prog. Ser. 50:73-81. Tracey, G., P.W. Johnson, R.W. Steele, P.E. Hargraves, and J. McN. Sieburth. 1988. A shift in photosynthetic picoplankton composition and its effect on bivalve mollusc nutrition; the 1985 "brown tide" in Narragansett Bay, Rhode Island. J. Shellfish Res. 7:671-675. Tracey, G., R. Steele, and L. Wright. 1990. Variable toxicity of the brown tide organism. Aureococcus anophagefferensin relation to environmental conditions for growth. , In: Toxic Marine Graneli, E., Sundstrdm, 8., Edler, L. & Anderson, D.M. (eds.), MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Phytoplankton. Elsevier, New York, pp. 233-237. White, W.A., T.R. Calnan, R.A. Morton, R.S. Kimble, T.G. Littleton, H.S. Nance, and K.E. Schmedes. 1983. Submerged lands of Texas Corpus Christi, TX.; Bureau of Economic Geology. Austin, TX: Univ. of Texas at Austin Bureau of Econ. Geol., 101 pp. MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Table 1. Gradient in Texas Estuaries. Listed from north to south; area at mean low tide (Diener, 1975), average annual precipitation (1951-1980; Larkin and Bomar, 1983), average annual freshwater inflow balance (1941-1976; Texas Department of Water Resources, 1982), and average annual commercial harvest (1962-1987; Texas Parks and Wildlife Department, 1988). Commercial Harvest Estuary Area Rainfall Inflow Finfish Shellfish (km 2 ) (cm y 1 ) (10 6 m 3 y 1 ) (10 3 kg y' 1 ) (10 3 kg y’) Sabine-Neches 183 142 16,107 5 332 Trinity-San Jacinto 1,416 112 12,284 190 4,060 Lavaca-Colorado 1,158 102 3,242 100 2,076 Guadalupe 551 91 2,545 80 1,545 Mission-Aransas 453 81 190 207 1,453 Nueces 433 76 509 151 544 Laguna Madre 1,139 69 -947 834 147 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Table 2. Location of sampling stations and sampling periods. Table gives estuary name, bay type, bay name, stations and years of the sampling. Estuary Bay Lavaca-Coiorado Secondary Secondary Primary Diversion Guadalupe Secondary Primary Nueces Secondary Primary Primary Laguna Madre Secondary Primary Name Lavaca Bay Lavaca Bay Matagorda Bay East Matagorda Upper San Antonio Bay Lower San Antonio Bay Nueces Bay Corpus Christi Bay Corpus Christi Bay Baffin Bay Laguna Madre Stations Period A 1984-1994 B 1988-1994 C, D 1988-1994 E. F 1993-1994 A, B 1987-1994 C. D 1987-1994 A, B 1988-1994 C. D 1988-1994 E 1990-1994 6. 24 1989-1993 189G, 189S 1989-1993 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Table 3. Lavaca-Colorado Estuary Mollusca distribution. Average numberin'2 for entire study period (1984-1994). Taxa A B c D E F Gastropoda Cuvier, 1797 Gastropoda (unidentified) 95 95 95 0 0 0 Ctenobranchia Schweigger, 1820 Hydrobiidae Troschel, 1857 Texadina sphinctostoma 189 189 0 0 0 0 (Abbott & Ladd, 1951) Caecidae Gray, 1850 Caecum pulchellum Stimpson, 1851 0 0 0 0 0 95 CaecumJohnson/' Winkley, 1908 0 0 165 95 95 0 Calyptraeidae Blainville, 1824 Crepidula fornicata (Linne, 1758) 0 0 0 284 0 0 Nassariidae Iredale, 1916 Nassarius acutus (Say, 1822) 189 158 113 189 189 95 Nassarius vibex (Say, 1822) 0 0 0 189 0 0 Columbellidae Swainson, 1840 Mitre!la lunata (Say, 1826) 0 0 95 0 0 0 Pleurobranchia Von Ihering, 1922 Acteocinidae Pilsbry, 1921 Acteocina candei(d’Orbigny, 1841) 321 662 95 189 591 284 Atyidae Thiele, 1926 Haminoea succinea (Conrad, 1846) 0 0 0 0 189 0 Entomotaeniata Cossman, 1896 Pyramidellidae Gray, 1840 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Odostomia sp. Fleming, 1813 284 189 0 0 0 0 Pyrgiscus sp. Philippi, 1841 0 142 138 95 0 0 EuHmastoma teres (Bush, 1885) 142 307 189 , 0 284 95 Eu/imostoma sp. Bartsch, 1916 0 0 0 0 189 0 Bivalvia Linne, 1758 =[Pelecypoda Goldfuss, 1820] Pelecypoda (unidentified) 158 142 118 701 0 95 Nuculoidea Dali, 1889 Nuculanidae H. & A. Adams, 1858 Nuculana acuta (Conrad, 1831) 95 0 95 252 725 0 Nuculana concentrica Say, 1824 142 284 221 142 95 0 Arcoidea Stoliczka, 1871 Arcidae Lamarck, 1809 Anadara ova/is (Bruguiere, 1879) 0 0 0 95 0 0 Mytiloidea Ferussac, 1822 Mytilidae Rafinesque, 1815 Brachidontes exustus (Linne, 1758) 95 95 0 0 0 95 Pterioidea Newell, 1965 Ostreidae Rafinesque, 1815 Crassostrea virginica (Gmelin, 1791) 0 95 0 0 0 0 Hippuritoidea Newell, 1965 Kelliidae Forbes & Flanley, 1848 AUgena texasiana Harry, 1969 0 0 95 95 0 0 Montacutidae Clark, 1855 Mysella planulata (Stimpson, 1851) 378 142 95 95 0 0 Mactridae Lamarck, 1809 Mulinia lateralis (Say, 1822) 714 544 1429 260 7162 993 Cultellidae Davis, 1935 Ensis minor Dali, 1900 350 0 0 0 0 0 Tellinidae Blainville, 1824 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Tellina texana Dali, 1900 0 0 0 95 0 0 Macoma tenta (Say, 1834) 0 0 0 95 0 0 Macoma mitchelli Dali, 1895 464 228 95 236 189 1059 Semelidae Stoliczka, 1870 Abra aequalis (Say, 1822) 0 0 0 176 0 0 Solecurtidae Orbigny, 1846 Tagelus plebeius (Lightfoot, 1786) 433 0 0 0 0 0 Veneridae Rafinesque, 1815 Mercenaria campechiensis 0 0 0 95 0 0 (Gmelin, 1791) Myoidea Stoliczka, 1870 Myidae Lamarck, 1809 Paramya subovata (Conrad, 1845) 0 0 0 700 0 0 Corbulidae Lamarck, 1818 Corbula contracta Say, 1822 0 0 0 819 0 0 Hiatellidae Gray, 1824 Hiatella arctica (Linne, 1767) 0 0 0 536 0 0 Pholadomyoidea Newell, 1965 Pandoridae Rafinesque, 1815 Pandora trilineata Say, 1822 0 95 0 95 0 0 Lyonsiidae Fischer, 1877 Lyonsia floridana Conrad, 1848 0 0 95 0 0 0 Periplomatidae Dali, 1895 Periploma cf. orbiculare Guppy, 1878 0 0 378 1179 0 0 Periploma margaritaceum 0 0 425 1182 0 0 (Lamarck, 1801) Scaphopoda Bronn, 1862 Dentaliidae Gray, 1834 Dentalium texasianum Philippi, 1849 0 0 0 95 0 0 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 2 Table 4. Guadalupe Estuary Mollusca distribution. Average number-nTfor entire study period (1987-1994). Taxa Gastropoda Cuvier, 1797 Gastropoda (unidentified) Ctenobranchia Schweigger, 1820 Hydrobiidae Troschel, 1857 Texadina sphinctostoma (Abbott & Ladd, 1951) Vitrinellidae Bush, 1897 Vitrinellid (unidentified) Caecidae Gray, 1850 Caecum pulchellum Stimpson, 1851 Caecum johnsoni Winkley, 1908 Calytraeidae Blainvilie, 1824 Crepidula fornicata (Linne, 1758) Nassariidae Iredale, 1916 Nassarius acutus (Say, 1822) Pleurobranchia Von Ihering, 1922 Acteocinidae Pilsbry, 1921 Acteocina candei(d’Orbigny, 1841) Entomotaeniata Cossmann, 1896 Pyramidellidae Gray, 1840 Odostomia sp. Fleming, 1813 Pyrgiscus sp. Philippi, 1841 Eu/imastoma teres (Bush, 1885) Eulimastoma sp. Bartsch, 1916 A 12478 14683 0 0 0 0 0 95 0 0 95 567 B C D 0 0 284 5138 4781 934 0 0 142 95 0 95 0 95 95 0 95 0 0 95 0 189 189 216 0 95 0 0 189 165 95 95 142 95 0 126 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Bivalvia Linne, 1758 =[Pelecypoda Goldfuss, 1820] Pelecypoda (unidentified) Nuculoidea Dali, 1889 Nuculanidae H. & A. Adams, 1858 Nuculana acuta (Conrad, 1831) Nuculana concentrica Say, 1824 Mytiioidea Ferussac, 1822 Mytilidae Rafinesque, 1815 Brachidontes exustus (Linne, 1758) Hippuritoidea Newell, 1965 Kelliidae Forbes & Hanley, 1848 AHgena texas/ana Harry, 1969 Montacutidae Clark, 1855 Mysella planulata (Stimpson, 1855) Mactridae Lamarck, 1809 Mulinia lateralis (Say, 1822) Rangia cuneata (Gray, 1831) Cultellidae Davis, 1935 Ensis minor Dali, 1900 Tellinidae Blainville, 1824 Tellina sp. Linne, 1758 Macoma tenta (Say, 1834) Macoma mitchelli Dali, 1895 Solecurtidae Orbigny, 1846 Tage/us p/ebeius (Lightfoot, 1786) Veneridae Rafinesque, 1815 Mercenaria campechiensis (Gmelin, 1791) Pholadomyoidea Newell, 1965 Pandoridae Rafinesque, 1815 0 0 0 0 0 0 6145 578 0 0 0 431 0 0 0 189 662 0 0 95 0 0 95 378 0 0 0 95 246 0 0 643 6802 2920 1642 142 0 0 95 95 473 95 0 95 0 0 95 441 244 490 0 189 189 0 95 95 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 27 Pandora trilineata Say, 1822 Lyonsiidae Fisher, 1877 Lyonsia floridana Conrad, 1848 Periplomatidae Dali, 1895 Periploma cf. orbiculare Guppy, Periploma margaritaceum (Lamarck, 1801) 1878 0 0 0 0 0 0 0 0 , 0 95 0 95 95 0 315 0 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 2 Table 5. Nueces Estuary Mollusca distribution. Average number-m'for entire study period (1988-1994). Taxa Moilusca Cuvier, 1795 Mollusca (unidentified) Gastropoda Cuvier, 1797 Gastropoda (unidentified) Ctenobranchia Schweigger, 1820 Vitrinellidae Bush, 1897 Vitrinella floridana Piisbry & McGinty, 1946 Vitrinellidae (unidentified) Caecidae Gray, 1850 Caecum johnsoni Winkley, 1908 Epitoniidae ss. Berry, 1910 Epitonium sp. Roding, 1798 Calyptraeidae Blainville, 1824 Crepidula plana Say, 1822 Nassariidae Iredale, 1916 Nassarius acutus (Say, 1822) Columbellidae Swainson, 1840 Anachis ostreiocola Sowerby, 1882 Pleurobranchia Von Ihering, 1922 Acteonidae Orbigny, 1835 A B c D E 0 0 0 95 95 0 0 0 95 0 0 0 0 95 0 0 0 662 0 0 0 236 425 95 0 0 0 95 0 0 0 95 95 0 0 0 189 95 95 0 0 95 0 0 Acteon punctostriatus 0 0 0 1135 473 (C.B. Adams, 1840) Acteocinidae Piisbry, 1921 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Acteocina candei (d’Orbigny, 1841) 95 236 0 113 0 Entomotaeniata Cassmann, 1896 Pyramidellidae Gray, 1840 Pyrgiscus sp. Philippi, 1841 0 O' 189 284 378 EuHmastoma teres (Bush, 1885) 0 95 0 0 0 EuHmastoma sp. Bartsch, 1916 95 0 0 236 0 Bivalvia Linne, 1758 =[Pelecypoda (Goldfuss, 1820)] Pelecypoda (unidentified) 0 0 189 189 0 Nuculoidea Dali, 1889 Nuculanidae H. & A. Adams, 1858 Nuculana acuta (Conrad, 1831) 0 95 851 284 252 Nuculana concentrica Say, 1824 0 0 473 0 0 Arcoidea Stoliczka, 1871 Arcidae Lamarck, 1809 Anadara transversa (Say, 1822) 0 0 0 0 95 Hippuritoidea Newell, 1965 Kelliidae Forbes & Hanley, 1848 AHgena texasiana Harry, 1969 0 95 0 605 378 Montacutidae Clark, 1855 Mysella planulata (Stimpson, 1851) 0 95 331 0 0 Crassatellidae Ferussac, 1821 Crassinel/a lunulata (Conrad, 1834) 0 0 189 0 0 Mactridae Lamarck, 1809 Mu/inia lateralis (Say, 1822) 1377 2902 697 1721 615 Solenidae Lamarck, 1809 Solen viridis Say, 1822 0 0 0 189 0 Cultellidae Davis, 1935 Ensis minor Dali, 1900 0 95 0 0 0 Tellinidae Blainville, 1824 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 30 Tellidora cris tata 0 0 0 0 95 H. & A. Adams, 1856 Macoma tenta (Say, 1834) 0 0 189 189 189 Macoma brevifrons (Say, 1834) 0 0 0 95 0 Macoma mitchelli Dali, 1895 1305 977 95 0 0 Semelidae Stoliczka, 1870 Abra aequa/is (Say, 1822) 0 0 95 0 189 Solecurtidae Orbigny, 1846 Tagelus divisus (Spengler, 1794) 0 0 284 0 0 Veneridae Rafinesque, 1815 Mercenaria campechiensis 0 0 0 95 0 (Grnelin, 1791) Pholadomyoidea Newell, 1965 Lyonsiidae Fischer, 1877 Lyonsia floridana Conrad, 1848 0 189 95 189 0 Periplomatidae Dali, 1895 Periploma cf. orbiculare Guppy, 1878 0 0 268 126 95 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Table 6. Laguna Madre-Baffin Bay Estuary Moliusca distribution. Average number-m'2 for entire study period (1989-1994). Taxa Gastropoda Cuvier, 1797 Gastropoda (unidentified) Ctenobranchia Schweigger, 1820 Littorinidae Gray, 1840 Littorina ziczac (Gmelin, 1791) Caecidae Gray, 1850 Caecum pulchellum Stimpson, 1851 Cerithiidae Fleming, 1822 Diastema varium (Pfieffer, 1840) Ceritheum lutosum (Menke, 1828) Calyptraeidae Blainville, 1824 Crepiduta fornicata (Linne, 1758) Nassariidae Iredale, 1916 Nassarius acutus (Say, 1822) Nassarius vibex (Say, 1822) Columbellidae Swainson, 1840 Anachis semip/icata Stearns, 1873 Anachis ostreico/a Sowerby 1882 Pleurobranchia Von Iheing, 1922 Acteonidae Orbigny, 1835 189G 221 0 2518 359 303 265 95 95 158 95 189S 6 24 95 0 95 95 0 0 1395 0 0 95 0 0 221 0 0 142 95 0 0 0 0 0 0 0 0 0 0 0 0 0 Rictaxis punctostriatus 0 0 529 511 (C.B. Adams, 1840) Acteocinidae Pilsbry, 1921 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Acteocina candei (d’Orbigny, 1841) 0 0 95 95 Atyidae Thiele, 1926 Haminoea anti!larum 5578 1418 0 0 (Orbigny, 1841) Entomotaeniata Cossman, 1896 Pyramidellidae Gray, 1840 Pyrgiscus sp. Philippi, 1841 340 425 0 0 Pyramidella crenulata 189 95 0 0 (Holmes, 1859) Sayel/a crosseana (Dali, 1885) 189 118 0 0 Boonea impressa (Say, 1822) 0 95 0 0 Bivalia Linne, 1758 =[Pelecypoda Goldfuss, 1820] Pelecypoda (unidentified) 95 0 0 0 Mytiloidea Ferussac, 1822 Mytilidae Rafinesque, 1815 Amygda/um papyria 265 95 0 0 (Conrad, 1846) Brachidontes exustus 520 95 0 0 (Linne, 1758) Hippuritoidea Newell, 1965 Cardiidae Lamarck, 1809 Laevicardium mortoni 126 95 0 0 (Conrad, 1830) Mactridae Lamarck, 1809 Mactra fragi/is Gmelin, 1791 359 95 0 0 Mulinia lateralis (Say, 1822) 95 315 352 1150 Tellinidae Blainville, 1824 TelUna sp. Linne, 1758 189 0 0 0 Tellina texana Dali, 1900 95 95 0 0 Tallina tampaensis Conrad, 1866 95 95 0 0 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA 33 Veneridae Rafinesque, 1815 Anomalocardiaauberiana 189 662 0 0 (Orbigny, 1842) Chione cancellata (Linne, 1767) 993 165 0 0 MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA Table 7. Summary of zoogeographic distributions and estuarine characteristics. Average salinity (PSU), number of species, abundance (n-rrT 2), and biomass (g-rrr 2) for each station over the entire study period. Estuary Station Salinity Species Abundance Biomass Lavaca-Colorado A 15 18 4,700 1.13 B 19 18 3,800 0.40 C 24 18 4,000 0.40 D 29 26 8,100 1.99 E 22 10 9,700 1.38 F 17 8 2,800 2.02 Guadalupe A 7 8 35,000 5.93 B 11 11 13,600 2.95 C 15 17 9,700 1.65 D 16 23 7,400 8.43 Nueces A 23 4 2,900 1.05 B 27 9 4,800 2.61 C 31 8 5,000 0.92 D 32 22 7,200 0.55 E 28 15 3,500 0.77 Laguna Madre 189G 38 24 13,000 10.41 189S 38 20 5,000 1.34 Baffin Bay 6 40 4 1,900 0.25 24 42 4 1,100 1.00 MONTAGNA AND KALKE; TEXAS INFAUNAL MOLLUSCA 2 Table 8. Dominant species in each estuary with overall ) in parentheses. average (n-m' Estuary 1st Dominant 2nd Dominant 3rd Dominant Lavaca- Mulinia lateralis Macoma mitchelli Acteocina cana/icu/ata Colorado (1900) (400) (400) Guadalupe Texadina sphinctostoma Mulinialateralis Macomamitchelli (6400) (4400) (400) Nueces Mulinia lateralis Macoma mitchelli Rictaxis punctostriatus (150) (50) (300) Laguna Haminoeaantillarum Caecum pulchellum Chinoe cance/lata Madre (400) (500) (600) Baffin Bay Mulinia lateralis Rictaxis punctostriatus Acteocina canaliculata (800) (500) (100) MONTAGNA AND KALKE: TEXAS INFAUNAL MOLLUSCA FIGURE LEGENDS Fig. 1. Location of south Texas estuaries and sampling stations. Fig 2. Bottom salinity values at each station during each sampling period in the Lavaca-Colorado Estuary. Fig 3. Bottom salinity values at each station during each sampling period in the Guadalupe Estuary. Fig 4. Bottom salinity values at each station during each sampling period in the Nueces Estuary. Fig 5. Bottom salinity values at each station during each sampling period in the Laguna Madre-Baffin Bay Estuary. Fig 6. Principal components analysis of molluscan communities at each station over the entire study period in the Lavaca-Colorado Estuary. Fig 7. Principal components analysis of molluscan communities at each station over the entire study period in the Guadalupe Estuary. Fig 8. Principal components analysis of molluscan communities at each station over the entire study period in the Nueces Estuary. Fig 9. Principal components analysis of molluscan communities at each station over the entire study period in the Laguna Madre-Baffin Bay Estuary. Fig 10. Mulinia lateralis average population abundance over entire estuary at each study period. A) In the Lavaca-Colorado (LC) and Guadalupe (GE) Estuaries. B) In the Nueces Estuary (NC) and Baffin Bay (BB). Lavaca-Colorado Estuary Guadalupe Estuary Nueces Estuary Laguna Madre-Baffin Bay Estuary Lavaca-Colorado Estuary Guadalupe Estuary Nueces Estuary Laguna Madre-Baffin Bay Estuary Mulinia lateralis RECENT BIOLOGICAL DATA COLLECTIONS Data taken from 1984-1992 can be found in the most recent comprehensive report (Montagna, P.A. 1992. Predicting long-term effects of freshwater inflow on macrobenthos in the Lavaca-Colorado and Guadalupe Estuaries. Year 2. Final Report to Texas Water Development Board. Technical Report No. TR/92-001, Marine Science Institute, The University of Texas, Port Aransas, TX, 105 pp.). The purpose of this data report is to transmit all data taken since that last report. HYDROGRAPHIC DATA Hydrographic measurements. Abbreviations: STA=Station, Z=Depth, SAL(R)=Salinity by refractometer, SAL(M)=Salinity by meter, COND=Conductivity, TEMP=Temperature, DO=dissolved oxygen, and ORP=oxidation redox potential. Missing values show with a period. Lavaca-Colorado Estuary Date STA z SAL(R) SAL(M) COND TEMP pH DO ORP 060CT92 A 0.00 22 19.6 31.70 25.28 8.99 8.25 0.186 060CT92 A 1.40 22 19.8 32.00 25.23 9.01 8.10 0.186 060CT92 B 0.00 25 23.0 36.50 24.66 8.56 7.73 0.190 060CT92 B 2.20 25 23.0 36.50 24.60 8.84 7.54 0.187 060CT92 C 0,00 28 25,9 40.40 24.52 8.46 7.35 0.187 060CT92 C 3.50 28 26.4 41.30 24.37 8.51 6.37 0.192 060CT92 D 0.00 31 27.0 42.10 24.27 8.18 6.80 0.201 060CT92 D 4.60 31 27.0 42.00 24.22 8.38 6.52 0.206 12JAN93 A 0.00 2 2.9 6.07 11.49 7.78 11.33 0.318 12JAN93 A 1.10 2 16.0 27.00 12.07 7.62 8.77 0.330 12JAN93 B 0.00 5 7.5 13.52 10.88 7.76 11.52 0.305 12JAN93 B 1.70 5 21.0 34.00 12.81 7.66 9.03 0.312 12JAN93 C 0.00 20 20.3 32.20 11.69 7.72 10.44 0.306 12JAN93 C 2.80 20 21.3 33.90 11.90 7.70 9.62 0.307 12JAN93 D 0.00 22 23.2 36.60 12.53 7.85 10.30 0.294 12JAN93 D 4.00 22 27.2 42.10 14.30 7.73 7.92 0.297 12JAN93 E 0.00 18 20.1 32.00 14.42 8.34 9.30 0.175 12JAN93 E 3.10 18 22.2 35.20 14.46 8.38 8.16 0.195 12JAN93 F 0.00 12 13.9 22.90 15.93 8.70 10.08 0.168 12JAN93 F 1.20 12 14.6 24.00 15.99 8.74 9.37 0.187 05APR93 A 0.00 0 0.0 0.69 17.41 7.98 10.16 0.191 05APR93 A 1.10 0 0.0 0.70 17.38 8.01 10.06 0.194 05APR93 B 0.00 8 8.3 14.74 17.82 7.77 9.74 0.266 05APR93 B 1.80 8 10.4 18.30 17.72 7.78 8.88 0.266 05APR93 C 0.00 15 15.2 25.50 18.84 7.76 9.30 0.267 05APR93 C 2.80 15 15.3 25.50 18.85 7.78 8.83 0.265 05APR93 D 0.00 0 20.0 31.90 19.16 7.91 9.05 0.257 05APR93 D 3.90 0 20.9 33.50 19.02 7.91 8.35 0.257 05APR93 E 0.00 16 15.6 26.00 19.69 7.92 9.60 0.258 05APR93 E 3.20 16 16.9 27.70 19.22 7.90 8.40 0.259 05APR93 F 0.00 4 1.6 4.06 18.63 8.23 10.66 0.241 05APR93 F 1.00 4 5.6 11.48 17.55 7.97 9.32 0.256 09JUL93 A 0.00 3 0.8 2.45 28.09 8.03 7.03 0.216 09JUL93 A 1.10 3 0.8 2.51 28.12 8.12 6.85 0.218 09JUL93 B 0.00 2 0.0 1.05 28.37 7.78 6.85 0.239 09JUL93 B 1.80 2 1.7 4.20 28.12 8.08 6.53 0.230 09JUL93 C 0.00 8 5.6 10.44 28.60 7.95 6.88 0.230 09JUL93 C 2.90 8 5.6 10.46 28.59 7.97 6.75 0.229 09JUL93 D 0.00 13 11.3 19.40 28.53 8.10 7.27 0.232 09JUL93 D 4.10 13 28.0 43.70 27.80 7.35 1.92 0.264 09JUL93 E 0.00 12 11.0 18.80 29.18 8.04 7.30 0.219 09JUL93 E 3.30 12 13.4 22.90 28.94 7.80 4.57 0.231 09JUL93 F 0.00 12 10.4 18.00 29.69 8.02 7.29 0.225 09JUL93 F 1.10 12 13.3 23.60 28.27 7.90 5.94 0.235 110CT93 A 0.00 18 16.6 27.10 23.37 8.06 7.05 0.230 110CT93 A 1.30 18 19.4 31.80 24.31 8.04 6.07 0.235 110CT93 B 0.00 22 20.4 32.80 23.74 8.03 7.64 0.241 110CT93 B 2.20 22 23.3 37.00 24.96 8.11 6.34 0.260 110CT93 C 0.00 28 26.1 40.90 25.06 8.10 7.03 0.262 110CT93 C 3.20 28 28.2 43.80 25.90 8.10 5.56 0.269 110CT93 D 0.00 28 26.9 42.00 26.11 8.03 7.23 0.224 110CT93 D 4.60 28 29.3 45.20 25.49 8.11 6.09 0.241 110CT93 E 0.00 32 29.3 45.10 25.64 8.18 6.41 0.228 110CT93 E 3.70 32 29.4 45.40 25.48 8.18 5.97 0.230 110CT93 F 0.00 28 26.1 40.90 24.90 8.20 7.50 0.232 110CT93 F 1.60 28 28.5 44.50 25.04 8.20 6.74 0.234 05JAN94 A 0.00 18 18.4 30.00 11.24 8.17 9.61 0.266 05JAN94 A 0.80 18 18.4 30.00 11.24 8.17 9.29 0.266 05JAN94 B 0.00 22 22.6 35.80 11.47 8.16 9.50 0.246 05JAN94 B 1.40 22 23.4 37.10 11.57 8.09 8.79 0.248 05JAN94 C 0.00 25 26.2 40.90 11.97 8.07 9.14 0.234 05JAN94 C 2.60 25 26.2 41.00 11.96 8.06 8.95 0.234 05JAN94 D 0.00 27 27.3 42.30 12.84 8.05 8.91 0.226 05JAN94 D 3.90 27 30.0 46.10 13.71 8.00 7.88 0.227 05JAN94 E 0.00 25 25.2 39.40 12.17 8.09 9.24 0.220 05JAN94 E 2.00 25 25.2 39.50 12.08 8.07 8.76 0.207 05JAN94 F 0.00 18 16.6 27.20 13.12 8.33 10.64 0.208 05JAN94 F 1.00 18 16.7 27.40 13.00 8.32 10.47 0.210 07APR94 A 0.00 15 14.0 23.80 15.61 7.66 8.88 0.091 07APR94 A 1.30 15 14.2 24.00 15.74 7.77 8.65 0.094 07APR94 B 0.00 20 20.1 33.50 16.73 7.38 9.29 0.085 07APR94 B 2.00 20 21.3 34.20 16.80 7.78 7.95 0.098 07APR94 C 0.00 25 24.9 39.00 17.60 7.71 7.77 0.127 07APR94 C 3.00 25 25.2 39.60 17.78 7.86 7.42 0.131 07APR94 D 0.00 26 25.9 40.50 17.81 7.74 8.05 0.149 07APR94 D 4.30 26 27.1 42.20 17.79 7.94 7.08 0.154 07APR94 E 0.00 26 25.4 39.80 17.95 7.68 8.55 0.159 07APR94 E 3.50 26 25.5 40.00 17.70 7.81 7.63 0.166 07APR94 F 0.00 22 19.7 32.10 17.32 7.68 9.75 0.193 07APR94 F 1.40 22 21.9 35.60 16.46 7.76 8.03 0.199 07JUL94 A 1.10 10 6.4 11.83 29.06 8.07 10.42 0.140 07JUL94 A 6.40 10 0.14 29.06 10.4 11.83 1.100 07JUL94 B 0.00 14 12.2 20.80 29.35 8.09 10.25 0.129 07JUL94 B 1.80 14 12.8 21.10 29.33 8.07 8.45 0.138 07JUL94 C 0.00 28 26.0 40.00 29.26 8.09 9.81 0.136 07JUL94 C 2.80 28 26.5 40.40 29.28 8.10 8.55 0.131 07JUL94 D 0.00 32 31.5 47.90 28.82 7.96 9.20 0.140 07JUL94 D 3.90 32 33.4 51.40 28.05 7.70 4.56 0.112 07JUL94 E 0.00 25 24.1 38.00 30.09 8.13 9.80 0.134 07JUL94 E 3.40 25 29.0 45.10 29.06 7.56 3.90 0.120 07JUL94 F 0.00 22 21.3 34.00 30.78 7.94 7.20 0.132 07JUL94 F 1.30 22 21.3 34.00 30.72 7.96 6.92 0.134 Guadalupe Estuary 070CT92 D 0.00 26 21.7 34.80 24.37 8.49 6.89 0.196 070CT92 D 1.40 26 21.8 34.80 24.38 8.59 6.46 0.198 070CT92 C 0.00 22 19.8 31.90 24.42 8.60 7.37 0.193 070CT92 C 1.90 22 19.8 31.90 24.43 8.71 6.75 0.195 070CT92 B 0.00 15 14.5 24.20 24.50 8.97 7.80 0.182 070CT92 B 1.50 15 15.0 25.00 24.53 8.94 6.95 0.174 070CT92 A 0.00 10 8.6 15.40 24.95 8.69 8.16 0.182 070CT92 A 1.20 10 8.7 15.10 24.93 8.97 7.63 0.183 12JAN93 A 0.00 10 10.2 17.10 11.79 8.11 14.20 0.260 12JAN93 A 1.20 10 18.4 29.60 12.43 8.07 8.64 0.275 12JAN93 B 0.00 12 12.6 20.90 12.13 8.27 12.60 0.273 12JAN93 B 1.70 12 19.4 31.10 11.68 8.13 9.67 0.283 12JAN93 C 0.00 15 15.2 24.90 12.20 8.40 12.70 0.269 12JAN93 C 1.90 15 21.0 33.60 11.92 8.09 9.46 0.285 12JAN93 D 0.00 22 22.0 35.00 12.37 8.02 10.79 0.284 12JAN93 D 1.60 22 25.0 39.20 12.01 7.95 9.84 0.291 05APR93 A 0.00 2 0.8 2.47 21.37 8.17 10.31 0.246 05APR93 A 0.90 2 3.5 7.86 19.52 7.71 8.55 0.266 05APR93 B 0.00 5 4.6 8.81 23.00 8.15 11.58 0.250 05APR93 B 1.50 5 5.9 10.76 18.91 7.84 8.25 0.260 05APR93 C 0.00 10 9.7 16.90 20.43 7.86 9.67 0.260 05APR93 C 1.70 10 9.8 17.10 18.96 7.60 7.66 0.270 05APR93 D 0.00 14 14.1 23.00 20.27 7.89 10.61 0.257 05APR93 D 1.30 14 15.8 26.10 19.23 7.74 8.45 0.262 09JUL93 D 0.00 2 0.0 0.77 29.95 7.76 7.43 0.227 09JUL93 D 1.20 2 0.0 0.92 29.79 7.89 6.95 0.228 09JUL93 C 0.00 4 1.1 2.91 29.98 8.02 7.40 0.217 09JUL93 C 1.70 4 4.1 8.35 29.17 7.95 6.35 0.228 09JUL93 B 0.00 1 0.0 0.98 30.17 7.86 6.95 0.230 09JUL93 B 1.50 1 0.0 0.95 30.02 7.89 6.64 0.234 09JUL93 A 0.00 1 0.0 0.60 30.39 7.69 6.76 0.232 09JUL93 A 1.10 1 0.0 0.60 30.41 7.80 6.54 0.234 110CT93 D 0.00 28 26.4 41.20 25.85 8.01 7.86 0.250 110CT93 D 2.00 28 26.4 41.30 25.81 8.17 7.52 0.257 110CT93 C 0.00 20 17.2 28.10 25.78 8.04 8.34 0.264 110CT93 C 2.20 20 17.6 28.60 25.76 8.17 7.86 0.264 110CT93 B 0.00 15 14.2 23.60 25.38 8.28 8.60 0.266 110CT93 B 2.20 15 14.4 26.20 25.51 8.22 8.31 0.270 110CT93 A 0.00 8 5.0 9.52 25.70 8.45 9.63 0.257 110CT93 A 1.70 8 5.0 9.52 25.69 8.47 9.31 0.261 05JAN94 D 0.00 20 19.8 31.90 14.01 8.27 11.26 0.209 05JAN94 D 0.90 20 19.8 31.90 13.97 8.29 10.24 0.210 05JAN94 C 0.00 20 19.4 31.40 13.90 8.39 10.69 0.208 05JAN94 C 1.30 20 19.4 31.40 13.82 8.37 10.31 0.209 05JAN94 B 0.00 19 17.8 29.10 13.54 8.59 13.20 0.202 05JAN94 B 1.20 19 17.8 29.10 13.51 8.57 13.05 0.203 05JAN94 A 0.00 10 8.7 15.30 14.26 8.28 10.50 0.211 05JAN94 A 0.70 10 8.7 15.30 14.19 8.27 10.30 0.212 07APR94 D 0.00 25 23.8 36.90 18.89 7.89 9.72 0.176 07APR94 D 1.40 25 24.0 37.00 18.93 7.96 9.05 0.178 07APR94 C 0.00 18 18.1 29.30 19.18 7.93 9.02 0.173 07APR94 C 2.00 18 18.2 29.40 18.77 7.94 7.82 0.175 07APR94 B 0.00 14 12.8 21.70 19.06 8.11 9.55 0.153 07APR94 B 1.60 14 12.8 21.70 19.07 8.17 9.47 0.155 07APR94 A 0.00 6 5.2 9.74 18.58 8.10 9.25 0.147 07APR94 A 1.50 6 5.2 9.76 18.60 8.18 9.19 0.151 07JUL94 D 0.00 10 7.5 13.30 30.29 8.31 11.84 0.107 07JUL94 D 1.20 10 7.5 13.37 30.25 8.44 11.77 0.109 07JUL94 C 0.00 22 20.8 32.60 30.28 8.00 10.65 0.132 07JUL94 C 1.60 22 21.2 ' 33.20 30.29 8.06 9.97 0.137 07JUL94 B 0.00 10 7.4 13.32 30.38 8.47 9.60 0.101 07JUL94 B 1.50 10 7.4 13.32 30.38 8.48 9.52 0.087 07JUL94 A 0.00 10 5.5 10.60 30.43 8.40 10.46 0.102 07JUL94 A 1.10 10 6.0 11.20 30.43 8.40 10.23 0.102 NUTRIENT CONCENTRATIONS Nutrient measurements take during sampling. Water depth is in m. Nutrient concentrations are in umol/l. Lavaca-Colorado Estuary Date Station Depth P0 4 SI0 4 N0 2 N0 3 NH 4 050CT92 A 0 1.089 134 .262 .329 .306 050CT92 A 1.4 1.096 132 .326 .409 .390 050CT92 B 0 1.010 118 .249 .353 .177 050CT92 B 2.2 1.526 157 .399 .407 .261 050CT92 C 0 .887 73. .164 .329 .048 050CT92 C 3.5 .970 73. .192 .326 .131 050CT92 D 0 .624 54. .150 .353 .084 050CT92 D 4.6 0 0 0 0 0 12JAN93 A 0 .493 135 .334 12.343 2.115 12JAN93 A 1.1 .678 39. .524 7.314 6.578 12JAN93 B 0 .429 84. .299 9.075 2.467 12JAN93 B 1.7 .275 43. .398 2.402 4.583 12JAN93 C 0 .262 45. .221 .153 .830 12JAN93 C 2.8 .237 28. .238 .187 .989 12JAN93 D 0 .397 .593 4.648 6.137 12JAN93 D 4. .230 .174 .165 1.145 12JAN93 E 0 .205 43. .274 1.540 1.729 12JAN93 E 3.1 .230 53. .325 2.281 2.671 12JAN93 F 0 .333 50. .480 .877 1.689 12JAN93 F 1.2 .256 70. .517 1.199 5.043 05APR93 A 0 .194 143 .372 19.178 1.440 05APR93 A 1.1 .183 146 .403 19.190 1.393 05APR93 B 0 .162 53. .525 7.107 2.368 05APR93 B 1.8 .151 44. .540 4.745 3.111 05APR93 C 0 .172 36. .111 .272 .557 05APR93 C 2.8 .151 21. .195 .063 1.579 05APR93 D 0 .226 33. .126 .008 .604 05APR93 D 3.9 .291 26. .325 .019 1.393 05APR93 E 0 .162 39. .149 .627 1.161 05APR93 E 3.2 .226 37. .240 .133 1.300 05APR93 F 0 .151 136 .654 37.494 2.229 05APR93 F 1.0 110CT93 A 0 .99 82. .93 .79 4.07 110CT93 A 1.3 1.13 65. 1.05 .79 3.12 110CT93 B 0 0.77 61. 0.67 .66 2.07 110CT93 B 2.2 1.14 65. 0.94 0.79 2.55 110CT93 C 0 0.71 53. 0.77 0.73 1.92 110CT93 C 3.2 0.83 25. 0.83 0.70 2.76 110CT93 D 0 0.59 5.4 0.73 0.70 2.33 110CT93 D 4.6 .70 9.9 0.78 0.68 3.25 110CT93 E 0 .76 4.2 0.69 0.65 2.18 110CT93 E 3.7 1.23 6.6 0.74 0.59 2.50 110CT93 F 0 0.80 48. 0.65 0.57 2.21 110CT93 F 1.6 0.93 54. 5.54 5.28 4.40 07APR94 A 0 .294 45. 1.018 2.079 1.704 07APR94 A 1.3 .392 45. 1.280 2.664 1.848 07APR94 B 0 .441 36. 1.242 4.010 5.232 07APR94 B 2.0 1.078 41. 1.805 3.835 7.200 07APR94 C 0 .686 18. .800 .625 4.656 07APR94 C 3.0 2.744 26. 1.894 0.000 11.520 07APR94 D 0 .392 17. .371 0.000 1.296 07APR94 D 4.3 .882 15. .730 0.000 3.600 07APR94 E 0 .441 17. .282 0.000 1.272 07APR94 E 3.5 .588 20. .422 0.000 1.872 07APR94 F 0 1.078 48. .384 4.747 6.240 07APR94 F 1.4 1.176 38. .768 .532 3.576 Guadalupe Estuary 070CT92 A 0 4.058 205 .359 .911 24.728 070CT92 A 1.2 3.437 196 .337 18.251 1.223 070CT92 B 0 1.791 178 .245 .673 .300 070CT92 B 1.5 2.026 178 .439 .625 .676 070CT92 C 0 1.539 172 .253 .616 .158 070CT92 C 1.9 1.384 156 .289 .665 .229 070CT92 D 0 .800 119 .204 .377 .169 070CT92 D 1.4 .882 116 .240 .486 .227 12JAN93 A 63.373 .68 2.070 28.80 12JAN93 A 0 .448 83. .700 1.519 12JAN93 B 0 .454 61. 14.371 13.887 1.088 12JAN93 B 1.7 .448 30. .556 7.365 1.900 12JAN93 C 0 .256 .296 .247 1.342 12JAN93 C 1.9 .384 .361 .356 2.479 12JAN93 D 0 .288 .247 .026 1.074 12JAN93 D 1.6 .333 .215 .046 1.038 05APR93 A 0 2.630 145 .959 98.602 3.111 05APR93 A 0.9 1.229 121 .790 80.975 5.851 05APR93 B 0 .647 117 20.510 19.835 9.288 05APR93 B 1.5 1.056 134 .783 25.383 1.393 05APR93 C 0 .151 68. .232 .577 4.180 05APR93 C 1.7 .172 55. .377 .752 8.220 05APR93 D 0 .172 40. .293 1.653 1.068 05APR93 D 1.3 .162 38. .247 .519 2.043 110CT93 A 0 3.32 135 18.17 17.83 2.13 110CT93 A 1.7 3.28 131 20.97 20.57 2.25 110CT93 B 0 3.14 115 0.60 0.60 2.40 110CT93 B 2.2 3.58 120 1.59 1.51 2.50 110CT93 C 0 1.95 81. 0.64 0.62 2.60 110CT93 D 0 1.81 81. 0.68 0.60 2.19 110CT93 D 2.0 1.59 73. 0.59 0.52 2.14 07APR94 A 0 4.312 127 1.472 71.64 1.536 07APR94 A 1.5 5.390 127 1.536 87.98 1.968 07APR94 B 0 1.862 94. 2.208 14.52 1.344 07APR94 B 1.5 2.058 99. 2.278 14.07 2.448 07APR94 C 0 .980 57. .947 4.507 3.192 07APR94 C 2.0 1.568 64. 1.523 4.702 6.336 07APR94 D 0 .441 41. 2.496 0.000 .960 07APR94 D 1.4 .372 41. .160 0.000 .792 BIOMASS DATA Biomass is measured for taxonomic groupings. Number and biomass per core, where biomass 2. a core is multiplied by 183 to get number or per m Lavaca-Colorado Estuary Date STA REP SEC Taxa n mg 060CT92 A 1 3 Crustacea 1 0.02 060CT92 A 1 3 Polychaeta 14 0.21 060CT92 A 1 10 Rhynchocoela 1 0.03 060CT92 A 1 10 Polychaeta 10 0.48 060CT92 A 2 3 Polychaeta 17 0.14 060CT92 A 2 10 Polychaeta 0 0 060CT92 A 3 3 Polychaeta 28 0.44 060CT92 A 3 10 Chironomid larvae 1 0.17 060CT92 A 3 10 Rhynchocoela 1 0.18 060CT92 A 3 10 Polychaeta 12 0.86 060CT92 B 1 3 Mollusca 1 0.04 060CT92 B 1 3 Polychaeta 19 0.44 060CT92 B 1 10 Polychaeta 6 0.4 060CT92 B 2 3 Polychaeta 19 0.38 060CT92 B 2 10 Polychaeta 4 1.07 060CT92 B 3 3 Polychaeta 3 0.16 060CT92 B 3 10 Polychaeta 2 0.25 060CT92 C 1 3 Mollusca 31 18.82 060CT92 C 1 3 Rhynchocoela 1 0.02 060CT92 C 1 3 Ophiuroidea 2 0.02 060CT92 C 1 3 Polychaeta 50 2.16 060CT92 C 1 10 Crustacea 1 0.05 060CT92 C 1 10 Rhynchocoela 1 0.03 060CT92 C 1 10 Polychaeta 24 1.9 060CT92 C 2 3 Mollusca 17 10.96 060CT92 C 2 3 Polychaeta 37 1.42 060CT92 C 2 10 Rhynchocoela 2 0.28 060CT92 C 2 10 Polychaeta 13 1.3 060CT92 C 3 3 Mollusca 22 13.12 060CT92 C 3 3 Rhynchocoela 2 0.03 060CT92 C 3 3 Polychaeta 27 0.74 060CT92 C 3 10 Rhynchocoela 2 0.02 060CT92 C 3 10 Polychaeta 11 1.38 060CT92 D 1 3 Mollusca 1 0.05 060CT92 D 1 3 Ophiuroidea 1 0.05 060CT92 D 1 3 Polychaeta 38 0.97 060CT92 D 1 10 Crustacea 3 1.66 060CT92 D 1 10 Mollusca 5 1.51 060CT92 D 1 10 Rhynchocoela 3 0.17 060CT92 D 1 10 Ophiuroidea 1 4.8 060CT92 D 1 10 Polychaeta 6 0.72 060CT92 D 2 3 Mollusca 2 0.28 060CT92 D 2 3 Polychaeta 58 1.54 060CT92 D 2 10 Mollusca 1 0.01 060CT92 D 2 10 Rhynchocoela 1 0.24 060CT92 D 2 10 Ophiuroidea 1 9.2 060CT92 D 2 10 Polychaeta 15 0.84 060CT92 D 3 3 Mollusca 1 0.1 060CT92 D 3 3 Polychaeta 38 1.56 060CT92 D 3 10 Mollusca 8 0.26 060CT92 D 3 10 Rhynchocoela 2 0.02 060CT92 D 3 10 Ophiuroidea 2 13.54 060CT92 D 3 10 Polychaeta 12 2.19 12JAN93 A 1 3 Chironomid larvae 1 0.07 12JAN93 A 1 3 Mollusca 4 0.28 12JAN93 A 1 3 Polychaeta 42 1.21 12JAN93 A 1 10 Polychaeta 1 0.1 12JAN93 A 2 3 Mollusca 3 0.21 12JAN93 A 2 3 Polychaeta 82 1.29 12JAN93 A 2 10 Mollusca 1 0.18 12JAN93 A 2 10 Polychaeta 3 1.4 12JAN93 A 3 3 Mollusca 3 0.2 12JAN93 A 3 3 Polychaeta 32 0.58 12JAN93 A 3 10 Polychaeta 0 0 12JAN93 B 1 3 Mollusca 3 0.15 12JAN93 B 1 3 Polychaeta 17 0.46 12JAN93 B 1 10 Polychaeta 2 0.06 12JAN93 B 2 3 Mollusca 3 0.11 12JAN93 B 2 3 Rhynchocoela 1 0.02 12JAN93 B 2 3 Polychaeta 39 0.58 12JAN93 B 2 10 Polychaeta 4 0.32 12JAN93 B 3 3 Crustacea 1 0.12 12JAN93 B 3 3 Mollusca 1 0.03 12JAN93 B 3 3 Polychaeta 33 0.85 12JAN93 B 3 10 Polychaeta 2 0.02 12JAN93 C 1 3 Crustacea 2 0.11 12JAN93 C 1 3 Mollusca 3 3.11 12JAN93 C 1 3 Rhynchocoela 1 0.08 12JAN93 C 1 3 Ophiuroidea 3 0.06 12JAN93 C 1 3 Polychaeta 85 0.74 12JAN93 C 1 10 Other 1 4.45 12JAN93 C 1 10 Polychaeta 18 3.88 12JAN93 C 2 3 Crustacea 2 0.06 12JAN93 C 2 3 Rhynchocoela 1 0.03 12JAN93 C 2 3 Polychaeta 51 2.51 12JAN93 C 2 10 Polychaeta 8 4.6 12JAN93 C 3 3 Mollusca 1 0.05 12JAN93 C 3 3 Rhynchocoela 2 0.04 12JAN93 C 3 3 Ophiuroidea 3 3.55 12JAN93 C 3 3 Polychaeta 49 1.24 12JAN93 C 3 3 Sipunculida 1 2.36 12JAN93 C 3 10 Rhynchocoela 2 0.26 12JAN93 C 3 10 Polychaeta 12 2.01 12JAN93 D 1 3 Crustacea 1 0.02 12JAN93 D 1 3 Mollusca 10 0.56 12JAN93 D 1 3 Rhynchocoela 3 0.05 12JAN93 D 1 3 Ophiuroidea 11 21.38 12JAN93 D 1 3 Polychaeta 55 5.14 12JAN93 D 1 10 Ophiuroidea 1 2.53 12JAN93 D 1 10 Polychaeta 4 1.05 12JAN93 D 2 3 Rhynchocoela 1 0.05 12JAN93 D 2 3 Polychaeta 13 2.63 12JAN93 D 2 10 Polychaeta 5 0.5 12JAN93 D 3 3 Crustacea 2 1.4 12JAN93 D 3 3 Rhynchocoela 2 0.04 12JAN93 D 3 3 Other 1 0.09 12JAN93 D 3 3 Ophiuroidea 1 0.2 12JAN93 D 3 3 Polychaeta 23 1.23 12JAN93 D 3 10 Polychaeta 6 6.77 12JAN93 E 1 3 Mollusca 2 0.06 12JAN93 E 1 3 Rhynchocoela 2 0.05 12JAN93 E 1 3 Ophiuroidea 1 0.02 12JAN93 E 1 3 Polychaeta 61 9.06 12JAN93 E 1 10 Rhynchocoela 1 0.03 12JAN93 E 1 10 Ophiuroidea 1 0.03 12JAN93 E 1 10 Polychaeta 8 4.91 12JAN93 E 2 3 Mollusca 2 0.1 12JAN93 E 2 3 Polychaeta 42 1.79 12JAN93 E 2 10 Polychaeta 12 17.5 12JAN93 E 3 3 Ophiuroidea 1 0.01 12JAN93 E 3 3 Polychaeta 31 3.65 12JAN93 E 3 10 Polychaeta 7 8.68 12JAN93 F 1 3 Mollusca 4 0.1 12JAN93 F 1 3 Polychaeta 26 2.01 12JAN93 F 1 10 Polychaeta 3 6.96 12JAN93 F 2 3 Mollusca 9 2.37 12JAN93 F 2 3 Mollusca 1 0.9 12JAN93 F 2 3 Other 2 0.39 12JAN93 F 2 3 Polychaeta 60 3.28 12JAN93 F 2 10 Rhynchocoela 2 7.94 12JAN93 F 2 10 Polychaeta 3 5.49 12JAN93 F 3 3 Mollusca 6 0.74 12JAN93 F 3 3 Polychaeta 38 1.76 12JAN93 F 3 10 Polychaeta 4 2.41 05APR93 A 1 3 Mollusca 1 0.22 05APR93 A 1 3 Rhynchocoela 1 0.05 05APR93 A 1 3 Other 1 0.06 05APR93 A 1 3 Polychaeta 48 0.92 05APR93 A 1 10 Mollusca 1 3.95 05APR93 A 1 10 Polychaeta 4 0.16 05APR93 A 2 3 Polychaeta 13 0.48 05APR93 A 2 10 Mollusca 1 2.43 05APR93 A 2 10 Polychaeta 1 0.04 05APR93 A 3 3 Rhynchocoela 5 0.19 05APR93 A 3 3 Polychaeta 43 0.74 05APR93 A 3 10 Mollusca 1 10.53 05APR93 A 3 10 Polychaeta 10 0.37 05APR93 B 1 3 Mollusca 7 4.64 05APR93 B 1 3 Polychaeta 60 2.44 05APR93 B 1 10 Mollusca 1 3.1 05APR93 B 1 10 Rhynchocoela 1 0.79 05APR93 B 1 10 Polychaeta 16 1.75 05APR93 B 2 3 Mollusca 2 3.96 05APR93 B 2 3 Rhynchocoela 1 1.93 05APR93 B 2 3 Polychaeta 41 3.6 05APR93 B 2 10 Rhynchocoela 1 7.02 05APR93 B 2 10 Other 4 0.45 05APR93 B 2 10 Polychaeta 8 0.49 05APR93 B 3 3 Polychaeta 11 3.86 05APR93 B 3 10 Rhynchocoela 2 0.58 05APR93 B 3 10 Polychaeta 9 10.22 05APR93 C 1 3 Crustacea 1 0.07 05APR93 C 1 3 Mollusca 33 10.19 05APR93 C 1 3 Polychaeta 4 0.71 05APR93 C 1 10 Mollusca 2 0.13 05APR93 C 1 10 Polychaeta 4 0.58 05APR93 C 2 3 Crustacea 5 0.15 05APR93 C 2 3 Mollusca 11 2.51 05APR93 C 2 3 Polychaeta 7 0.6 05APR93 C 2 10 Mollusca 1 0.1 05APR93 C 2 10 Polychaeta 8 5.56 05APR93 C 3 3 Crustacea 2 0.09 05APR93 C 3 3 Mollusca 10 5.18 05APR93 C 3 3 Polychaeta 3 0.65 05APR93 C 3 10 Polychaeta 4 1.41 05APR93 D 1 3 Crustacea 1 0.02 05APR93 D 1 3 Mollusca 1 0.02 05APR93 D 1 3 Rhynchocoela 1 0.04 05APR93 D 1 3 Other 1 0.12 05APR93 D 1 3 Polychaeta 85 3.96 05APR93 D 1 10 Rhynchocoela 1 0.74 05APR93 D 1 10 Ophiuroidea 4 25.38 05APR93 D 1 10 Polychaeta 12 9.5 05APR93 D 2 3 Rhynchocoela 1 0.23 05APR93 D 2 3 Other 2 0.54 05APR93 D 2 3 Ophiuroidea 1 0.1 05APR93 D 2 3 Polychaeta 72 2.81 05APR93 D 2 3 Sipunculida 1 0.06 05APR93 D 2 10 Rhynchocoela 4 13.58 05APR93 D 2 10 Polychaeta 7 4.2 05APR93 D 3 3 Crustacea 1 0.02 05APR93 D 3 3 Mollusca 1 0.04 05APR93 D 3 3 Polychaeta 51 1.56 05APR93 D 3 10 Other 1 183.6 05APR93 D 3 10 Polychaeta 12 7.36 05APR93 E 1 3 Crustacea 1 0.04 05APR93 E 1 3 Mollusca 2 1.25 05APR93 E 1 3 Rhynchocoela 3 0.57 05APR93 E 1 3 Polychaeta 37 2.11 05APR93 E 1 10 Polychaeta 5 2.54 05APR93 E 2 3 Mollusca 5 1.49 05APR93 E 2 3 Rhynchocoela 1 0.22 05APR93 E 2 3 Polychaeta 34 2.33 05APR93 E 2 10 Other 1 0.98 05APR93 E 2 10 Polychaeta 6 0.41 05APR93 E 3 3 Crustacea 1 0.07 05APR93 E 3 3 Mollusca 5 2.13 05APR93 E 3 3 Ophiuroidea 1 0.34 05APR93 E 3 3 Polychaeta 44 5.81 05APR93 E 3 10 Rhynchocoela 3 0.41 05APR93 E 3 10 Polychaeta 8 3.6 05APR93 F 1 3 Mollusca 12 6.72 05APR93 F 1 3 Other 4 0.07 05APR93 F 1 3 Polychaeta 47 1.78 05APR93 F 1 10 Mollusca 2 29.29 05APR93 F 1 10 Rhynchocoela 2 1.7 05APR93 F 1 10 Other 2 0.03 05APR93 F 1 10 Polychaeta 12 8.77 05APR93 F 2 3 Mollusca 8 3.68 05APR93 F 2 3 Rhynchocoela 1 0.12 05APR93 F 2 3 Polychaeta 46 1.76 05APR93 F 2 10 Rhynchocoela 3 4.8 05APR93 F 2 10 Polychaeta 9 5.28 05APR93 F 3 3 Mollusca 7 3.02 05APR93 F 3 3 Polychaeta 77 3.05 05APR93 F 3 10 Mollusca 2 20.48 05APR93 F 3 10 Rhynchocoela 1 0.01 05APR93 F 3 10 Polychaeta 4 5.76 09JUL93 A 1 3 Mollusca 3 0.17 09JUL93 A 1 3 Polychaeta 15 0.2 09JUL93 A 1 10 Chironomid larvae 1 0.08 09JUL93 A 2 3 Chironomid larvae 1 0.06 09JUL93 A 2 3 Mollusca 5 0.62 09JUL93 A 2 3 Polychaeta 25 0.36 09JUL93 A 2 10 Mollusca 1 5.92 09JUL93 A 2 10 Polychaeta 2 0.03 09JUL93 A 3 3 Mollusca 1 0.12 09JUL93 A 3 3 Polychaeta 21 0.17 09JUL93 A 3 10 Mollusca 1 6.75 09JUL93 A 3 10 Polychaeta 4 0.16 09JUL93 B 1 3 Chironomid larvae 1 0.07 09JUL93 B 1 3 Polychaeta 4 0.3 09JUL93 B 1 10 Polychaeta 2 0.44 09JUL93 B 2 3 Mollusca 3 0.2 09JUL93 B 2 10 Polychaeta 7 1.28 09JUL93 B 3 3 Polychaeta 6 0.28 09JUL93 B 3 10 Polychaeta 10 1.1 09JUL93 C 1 3 Polychaeta 12 0.73 09JUL93 C 1 10 Polychaeta 1 0.34 09JUL93 C 2 3 Mollusca 2 0.98 09JUL93 C 2 3 Polychaeta 11 0.53 09JUL93 C 2 10 Polychaeta 5 0.73 09JUL93 C 3 3 Mollusca 2 1.72 09JUL93 C 3 3 Polychaeta 22 1.12 09JUL93 C 3 10 Mollusca 4 0.53 09JUL93 C 3 10 Rhynchocoela 1 0.08 09JUL93 C 3 10 Polychaeta 5 9.74 09JUL93 D 1 3 Crustacea 1 0.02 09JUL93 D 1 3 Mollusca 1 0.02 09JUL93 D 1 3 Other 1 0.12 09JUL93 D 1 3 Polychaeta 7 0.48 09JUL93 D 1 10 Crustacea 2 1.98 09JUL93 D 1 10 Mollusca 1 0.08 09JUL93 D 1 10 Polychaeta 5 2.09 09JUL93 D 2 3 Crustacea 4 0.03 09JUL93 D 2 3 Polychaeta 3 0.11 09JUL93 D 2 10 Crustacea 1 0.54 09JUL93 D 2 10 Mollusca 2 0.06 09JUL93 D 2 10 Rhynchocoela 1 0.26 09JUL93 D 2 10 Polychaeta 7 0.29 09JUL93 D 3 3 Mollusca 1 0.57 09JUL93 D 3 3 Polychaeta 15 0.79 09JUL93 D 3 10 Mollusca 3 0.31 09JUL93 D 3 10 Rhynchocoela 1 0.72 09JUL93 D 3 10 Ophiuroidea 1 4.66 09JUL93 D 3 10 Polychaeta 5 2.03 09JUL93 E 1 3 Mollusca 11 0.28 09JUL93 E 1 3 Polychaeta 10 2.33 09JUL93 E 1 10 Polychaeta 5 0.62 09JUL93 E 2 3 Mollusca 14 2.68 09JUL93 E 2 3 Polychaeta 11 0.24 09JUL93 E 2 10 Polychaeta 7 4 09JUL93 E 3 3 Mollusca 5 0.71 09JUL93 E 3 3 Rhynchocoela 1 0.04 09JUL93 E 3 3 Polychaeta 8 0.19 09JUL93 E 3 10 Polychaeta 7 3.26 09JUL93 F 1 3 Rhynchocoela 1 0.23 09JUL93 F 1 3 Polychaeta 15 1.69 09JUL93 F 1 10 Polychaeta 19 4.56 09JUL93 F 2 3 Polychaeta 7 1.45 09JUL93 F 2 10 Polychaeta 20 5.82 09JUL93 F 3 3 Mollusca 1 0.02 09JUL93 F 3 3 Polychaeta 26 2.58 09JUL93 F 3 10 Polychaeta 14 2.68 110CT93 A 1 3 Polychaeta 5 0.13 110CT93 A 1 10 Polychaeta 0 0 110CT93 A 2 3 Polychaeta 7 0.22 110CT93 A 2 10 Polychaeta 0 0 110CT93 A 3 3 Polychaeta 0 0 110CT93 A 3 10 Polychaeta 0 0 110CT93 B 1 3 Mollusca 1 0.01 110CT93 B 1 3 Polychaeta 1 0.01 110CT93 B 1 10 Polychaeta 3 0.87 110CT93 B 2 3 Rhynchocoela 2 0.04 110CT93 B 2 3 Polychaeta 3 0.24 110CT93 B 2 10 Polychaeta 6 1.29 110CT93 B 3 3 Polychaeta 1 0.11 110CT93 B 3 10 Polychaeta 3 0.28 110CT93 C 1 3 Rhynchocoela 1 0.13 110CT93 C 1 3 Ophiuroidea 1 0.02 110CT93 C 1 3 Polychaeta 16 1.27 110CT93 C 1 10 Polychaeta 12 1.44 110CT93 C 2 3 Mollusca 2 0.04 110CT93 C 2 3 Ophiuroidea 2 0.04 110CT93 C 2 3 Polychaeta 23 1.7 110CT93 C 2 10 Rhynchocoela 1 0.07 110CT93 C 2 10 Polychaeta 17 2.79 110CT93 C 3 3 Mollusca 2 0.22 110CT93 C 3 3 Other 1 0.69 110CT93 C 3 3 Ophiuroidea 1 0.01 1100T93 C 3 3 Polychaeta 6 1.47 110CT93 C 3 10 Polychaeta 2 1.12 110CT93 D 1 3 Rhynchocoela 1 0.01 110CT93 D 1 3 Polychaeta 8 0.79 1100T93 D 1 10 Crustacea 1 0.02 110CT93 D 1 10 Rhynchocoela 2 0.71 110CT93 D 1 10 Ophiuroidea 1 18.51 110CT93 D 1 10 Polychaeta 10 2.75 110CT93 D 2 3 Mollusca 4 0.18 110CT93 D 2 3 Rhynchocoela 2 0.72 110CT93 D 2 3 Ophiuroidea 1 11.6 110CT93 D 2 3 Polychaeta 10 0.22 110CT93 D 2 10 Mollusca 3 0.26 110CT93 D 2 10 Rhynchocoela 2 0.03 110CT93 D 2 10 Ophiuroidea 1 6.84 110CT93 D 2 10 Polychaeta 6 0.31 110CT93 D 3 3 Mollusca 5 0.3 110CT93 D 3 3 Polychaeta 4 0.18 110CT93 D 3 10 Mollusca 2 0.04 110CT93 D 3 10 Rhynchocoela 1 0.01 110CT93 D 3 10 Ophiuroidea 1 2.22 110CT93 D 3 10 Polychaeta 8 0.2 110CT93 E 1 3 Mollusca 2 0.75 110CT93 E 1 3 Polychaeta 10 0.33 110CT93 E 1 10 Polychaeta 9 4.41 110CT93 E 2 3 Mollusca 1 0.07 110CT93 E 2 3 Polychaeta 13 0.51 110CT93 E 2 10 Mollusca 1 0.07 110CT93 E 2 10 Polychaeta 12 4.33 110CT93 E 3 3 Mollusca 1 0.11 110CT93 E 3 3 Polychaeta 7 0.24 110CT93 E 3 10 Polychaeta 7 6.35 110CT93 F 1 3 Crustacea 4 0.36 110CT93 F 1 3 Polychaeta 6 0.46 110CT93 F 1 10 Polychaeta 5 3.84 110CT93 F 2 3 Crustacea 3 0.33 110CT93 F 2 3 Rhynchocoela 1 0.01 110CT93 F 2 3 Polychaeta 12 0.35 110CT93 F 2 10 Polychaeta 3 0.94 110CT93 F 3 3 Crustacea 1 0.09 11GCT93 F 3 3 Mollusca 1 0.05 110CT93 F 3 3 Rhynchocoela 1 0.01 11QCT93 F 3 3 Polychaeta 9 0.45 110CT93 F 3 10 Polychaeta 0 0 05JAN94 A 1 3 Crustacea 1 0.01 05JAN94 A 1 3 Mollusca 3 0.47 05JAN94 A 1 3 Polychaeta 29 0.93 05JAN94 A 1 10 Polychaeta 1 0.02 05JAN94 A 2 3 Crustacea 3 0.07 05JAN94 A 2 3 Mollusca 5 0.16 05JAN94 A 2 3 Rhynchocoela 1 0.39 05JAN94 A 2 3 Polychaeta 35 1.68 05JAN94 A 2 10 Polychaeta 2 0.02 05JAN94 A 3 3 Mollusca 11 0.88 05JAN94 A 3 3 Polychaeta 23 0.36 05JAN94 A 3 10 Polychaeta 0 0 05JAN94 B 1 3 Mollusca 7 0.11 05JAN94 B 1 3 Polychaeta 7 0.21 05JAN94 B 1 10 Crustacea 1 4.89 05JAN94 B 1 10 Polychaeta 5 5.23 05JAN94 B 2 3 Mollusca 6 1.74 05JAN94 B 2 3 Polychaeta 11 0.84 05JAN94 B 2 10 Polychaeta 1 0.25 05JAN94 B 3 3 Mollusca 7 0.18 05JAN94 B 3 3 Polychaeta 13 1.65 05JAN94 B 3 10 Mollusca 2 6.65 05JAN94 B 3 10 Polychaeta 4 24.81 05JAN94 C 1 3 Crustacea 1 0.01 05JAN94 C 1 3 Polychaeta 11 2.67 05JAN94 C 1 10 Rhynchocoela 1 0.31 05JAN94 C 1 10 Polychaeta 3 5.78 05JAN94 C 2 3 Mollusca 1 0.61 05JAN94 C 2 3 Rhynchocoela 2 0.45 05JAN94 C 2 3 Ophiuroidea 1 0.01 05JAN94 C 2 3 Polychaeta 26 1.8 05JAN94 C 2 10 Rhynchocoela 1 0.35 05JAN94 C 2 10 Polychaeta 6 1.32 05JAN94 C 3 3 Rhynchocoela 2 0.13 05JAN94 C 3 3 Polychaeta 21 2.98 05JAN94 C 3 10 Polychaeta 4 0.54 05JAN94 D 1 3 Crustacea 2 1.67 05JAN94 D 1 3 Mollusca 2 0.16 05JAN94 D 1 3 Rhynchocoela 1 0.01 05JAN94 D 1 3 Ophiuroidea 1 0.97 05JAN94 D 1 3 Polychaeta 21 5.73 05JAN94 D 1 10 Ophiuroidea 1 1.68 05JAN94 D 2 3 Crustacea 4 0.05 05JAN94 D 2 3 Other 3 0.1 05JAN94 D 2 3 Ophiuroidea 1 1.19 05JAN94 D 2 3 Polychaeta 21 1.61 05JAN94 D 2 10 Mollusca 1 0.05 05JAN94 D 2 10 Rhynchocoela 2 0.48 05JAN94 D 2 10 Ophiuroidea 1 4.54 05JAN94 D 2 10 Polychaeta 18 4.61 05JAN94 D 3 3 Crustacea 1 0.01 05JAN94 D 3 3 Mollusca 1 0.01 05JAN94 D 3 3 Rhynchocoela 2 0.15 05JAN94 D 3 3 Other 4 0.46 05JAN94 D 3 3 Ophiuroidea 1 0.03 05JAN94 D 3 3 Polychaeta 26 1.58 05JAN94 D 3 10 Crustacea 4 4.1 05JAN94 D 3 10 Mollusca 7 0.2 05JAN94 D 3 10 Rhynchocoela 2 4.15 05JAN94 D 3 10 Ophiuroidea 1 2.76 05JAN94 D 3 10 Polychaeta 21 5.28 05JAN94 E 1 3 Mollusca 73 11.45 05JAN94 E 1 3 Ophiuroidea 1 0.02 05JAN94 E 1 3 Polychaeta 41 3.54 05JAN94 E 1 10 Polychaeta 23 13.64 05JAN94 E 2 3 Crustacea 2 0.24 05JAN94 E 2 3 Mollusca 76 8.81 05JAN94 E 2 3 Other 1 0.04 05JAN94 E 2 3 Polychaeta 17 1.44 05JAN94 E 2 10 Mollusca 1 0.09 05JAN94 E 2 10 Rhynchocoela 1 0.07 05JAN94 E 2 10 Polychaeta 26 9.29 05JAN94 E 3 3 Crustacea 1 0.02 05JAN94 E 3 3 Mollusca 106 8.17 05JAN94 E 3 3 Other 2 0.13 05JAN94 E 3 3 Ophiuroidea 1 0.01 05JAN94 E 3 3 Polychaeta 22 2.29 05JAN94 E 3 10 Mollusca 2 2.71 05JAN94 E 3 10 Polychaeta 36 8.09 05JAN94 F 1 3 Crustacea 1 5.64 05JAN94 F 1 3 Mollusca 10 7.99 05JAN94 F 1 3 Other 1 0.46 05JAN94 F 1 3 Polychaeta 12 0.32 05JAN94 F 1 10 Other 1 0.79 05JAN94 F 2 3 Crustacea 1 0.13 05JAN94 F 2 3 Mollusca 13 3.24 05JAN94 F 2 3 Rhynchocoela 1 0.18 05JAN94 F 2 3 Polychaeta 16 4.65 05JAN94 F 2 10 Polychaeta 1 3.99 05JAN94 F 3 3 Mollusca 16 4.97 05JAN94 F 3 3 Polychaeta 14 1.27 05JAN94 F 3 10 Mollusca 7 4.37 05JAN94 F 3 10 Rhynchocoela 1 0.07 05JAN94 F 3 10 Polychaeta 3 0.68 07APR94 A 1 3 Crustacea 1 0.14 07APR94 A 1 3 Mollusca 8 2.4 07APR94 A 1 3 Polychaeta 24 1.12 07APR94 A 1 10 Rhynchocoela 1 0.87 07APR94 A 1 10 Polychaeta 7 1.44 07APR94 A 2 3 Crustacea 2 0.02 07APR94 A 2 3 Mollusca 9 0.82 07APR94 A 2 3 Polychaeta 25 2.21 07APR94 A 2 10 Crustacea 1 0.01 07APR94 A 2 10 Mollusca 1 12.18 07APR94 A 2 10 Polychaeta 14 1.93 07APR94 ,­A 3 3 Crustacea 1 0.05 07APR94 A 3 3 Mollusca 10 3.54 07APR94 A 3 3 Polychaeta 38 1.85 07APR94 A 3 10 Mollusca 1 3.06 07APR94 A 3 10 Polychaeta 19 4.35 07APR94 B 1 3 Other 1 0.13 07APR94 B 1 3 Polychaeta 31 3.14 07APR94 B 1 10 Mollusca 1 8.36 07APR94 B 1 10 Polychaeta 6 4.45 07APR94 B 2 3 Mollusca 3 1.03 07APR94 B 2 3 Polychaeta 28 1.62 07APR94 B 2 10 Rhynchocoela 1 0.18 07APR94 B 2 10 Polychaeta 3 5.03 07APR94 B 3 3 Mollusca 4 0.32 07APR94 B 3 3 Rhynchocoela 1 0.49 07APR94 B 3 3 Polychaeta 13 0.72 07APR94 B 3 10 Rhynchocoela 1 0.05 07APR94 B 3 10 Polychaeta 4 1.49 07APR94 C 1 3 Rhynchocoela 1 0.39 07APR94 C 1 3 Polychaeta 6 2.06 07APR94 C 1 10 Polychaeta 3 0.37 07APR94 C 2 3 Polychaeta 4 1.57 07APR94 C 2 10 Polychaeta 7 1.23 07APR94 C 3 3 Polychaeta 4 0.73 07APR94 C 3 10 Polychaeta 6 1.13 07APR94 D 1 3 Crustacea 3 0.07 07APR94 D 1 3 Rhynchocoela 1 0.07 07APR94 D 1 3 Other 2 0.2 07APR94 D 1 3 Polychaeta 18 0.81 07APR94 D 1 10 Rhynchocoela 2 2.91 07APR94 D 1 10 Ophiuroidea 1 6.19 07APR94 D 1 10 Polychaeta 29 5.47 07APR94 D 2 3 Other 1 0.05 07APR94 D 2 3 Polychaeta 10 0.94 07APR94 D 2 10 Ophiuroidea 1 2.08 07APR94 D 2 10 Polychaeta 13 17.16 07APR94 D 3 3 Crustacea 1 0.86 07APR94 D 3 3 Mollusca 2 0.91 07APR94 D 3 3 Rhynchocoela 1 0.2 07APR94 D 3 3 Ophiuroidea 1 3.21 07APR94 D 3 3 Polychaeta 17 0.79 07APR94 D 3 10 Crustacea 1 0.01 07APR94 D 3 10 Mollusca 2 0.05 07APR94 D 3 10 Ophiuroidea 1 3.36 07APR94 D 3 10 Polychaeta 12 18.22 07APR94 E 1 3 Mollusca 30 11.55 07APR94 E 1 3 Ophiuroidea 1 0.32 07APR94 E 1 3 Polychaeta 18 0.49 07APR94 E 1 10 Molluscs 1 0.72 07APR94 E 1 10 Ophiuroidea 2 0.11 07APR94 E 1 10 Polychaeta 21 10.83 07APR94 E 2 3 Crustacea 4 0.07 07APR94 E 2 3 Mollusca 1 16.1 07APR94 E 2 3 Polychaeta 26 1.83 07APR94 E 2 10 Mollusca 1 0.66 07APR94 E 2 10 Rhynchocoela 1 0.35 07APR94 E 2 10 Polychaeta 25 13.12 07APR94 E 3 3 Crustacea 6 0.7 07APR94 E 3 3 Mollusca 34 17.77 07APR94 E 3 3 Other 1 0.04 07APR94 E 3 3 Ophiuroidea 1 0.13 07APR94 E 3 3 Polychaeta 34 1.13 07APR94 E 3 10 Polychaeta 25 9.01 07APR94 F 1 3 Crustacea 1 0.02 07APR94 F 1 3 Mollusca 4 5.28 07APR94 F 1 3 Polychaeta 8 0.65 07APR94 F 1 10 Mollusca 1 4.94 07APR94 F 2 3 Crustacea 2 0.01 07APR94 F 2 3 Mollusca 4 2.65 07APR94 F 2 3 Polychaeta 9 0.55 07APR94 F 2 10 Mollusca 3 26.55 07APR94 F 2 10 Polychaeta 5 6.89 07APR94 F 3 3 Crustacea 5 0.13 07APR94 F 3 3 Mollusca 3 1.08 07APR94 F 3 3 Polychaeta 5 3.35 07APR94 F 3 10 Polychaeta 4 4.94 Guadalupe Estuary 070CT92 A 1 3 Chironomid larvae 1 0.11 070CT92 A 1 3 Mollusca 82 41.52 070CT92 A 1 3 Polychaeta 2 0.19 070CT92 A 1 10 Mollusca 1 0.2 070CT92 A 2 3 Chironomid larvae 2 0.3 070CT92 A 2 3 Mollusca 15 9.16 070CT92 A 2 3 Polychaeta 2 0.13 070CT92 A 2 10 Chironomid larvae 1 0.19 070CT92 A 2 10 Polychaeta 3 0.79 070CT92 A 3 3 Mollusca 116 78.81 070CT92 A 3 3 Polychaeta 4 0.64 070CT92 A 3 10 Polychaeta 0 0 070CT92 B 1 3 Mollusca 1 0.11 070CT92 B 1 3 Polychaeta 83 3.75 070CT92 B 1 10 Polychaeta 23 8.37 070CT92 B 2 3 Polychaeta 74 3.86 070CT92 B 2 10 Chironomid larvae 1 0.16 070CT92 B 2 10 Rhynchocoela 1 0.85 070CT92 B 2 10 Polychaeta 24 4.76 070CT92 B 3 3 Chironomid larvae 1 0.03 070CT92 B 3 3 Mollusca 3 6.82 070CT92 B 3 *3 Polychaeta 109 5.44 070CT92 B 3 10 Rhynchocoela 1 1.54 070CT92 B 3 10 Polychaeta 22 5.69 070CT92 C 1 3 Polychaeta 27 1.42 070CT92 C 1 10 Crustacea 1 1.2 070CT92 C 1 10 Rhynchocoela 2 0.26 070CT92 C 1 10 Polychaeta 4 0.63 070CT92 C 2 3 Polychaeta 18 1.2 070CT92 C 2 10 Crustacea 1 0.24 070CT92 C 2 10 Rhynchocoela 5 0.69 070CT92 C 2 10 Polychaeta 16 2.32 070CT92 C 3 3 Mollusca 1 0.02 070CT92 C 3 3 Rhynchocoela 2 0.63 070CT92 C 3 3 Polychaeta 29 1.62 070CT92 C 3 10 Rhynchocoela 2 0.47 070CT92 C 3 10 Polychaeta 7 0.95 070CT92 D 1 3 Crustacea 1 0.15 070CT92 D 1 3 Mollusca 1 4.61 070CT92 D 1 3 Polychaeta 7 0.57 070CT92 D 1 10 Polychaeta 3 0.59 070CT92 D 2 3 Crustacea 1 0.02 070CT92 D 2 3 Mollusca 1 0.12 070CT92 D 2 3 Rhynchocoela 2 2.53 070CT92 D 2 3 Polychaeta 12 0.85 070CT92 D 2 10 Other 1 0.07 070CT92 D 2 10 Polychaeta 3 0.15 070CT92 D 3 3 Mollusca 1 0.14 070CT92 D 3 3 Rhynchocoela 1 0.02 070CT92 D 3 3 Polychaeta 8 0.33 070CT92 D 3 10 Polychaeta 3 0.76 12JAN93 A 1 3 Mollusca 138 31.84 12JAN93 A 1 3 Rhynchocoela 2 0.12 12JAN93 A 1 3 Polychaeta 6 0.18 12JAN93 A 1 10 Chironomid larvae 1 0.57 12JAN93 A 1 10 Polychaeta 1 0.58 12JAN93 A 2 3 Mollusca 156 34.69 12JAN93 A 2 3 Rhynchocoela 1 0.06 12JAN93 A 2 3 Polychaeta 10 0.38 12JAN93 A 2 10 Rhynchocoela 1 0.15 12JAN93 A 2 10 Polychaeta 1 0.06 12JAN93 A 3 3 Crustacea 1 0.01 12JAN93 A 3 3 Mollusca 134 29.76 12JAN93 A 3 3 Rhynchocoela 1 0.01 12JAN93 A 3 3 Polychaeta 11 0.43 12JAN93 A 3 10 Polychaeta 1 0.16 12JAN93 B 1 3 Mollusca 55 5.7 12JAN93 B 1 3 Rhynchocoela 1 0.01 12JAN93 B 1 3 Polychaeta 33 4.05 12JAN93 B 1 10 Polychaeta 5 2.31 12JAN93 B 2 3 Mollusca 87 3.43 12JAN93 B 2 3 Polychaeta 41 2.74 12JAN93 B 2 10 Mollusca 1 0.31 12JAN93 B 2 10 Polychaeta 1 0.1 12JAN93 B 3 3 Mollusca 63 0.96 12JAN93 B 3 3 Rhynchocoela 1 0.05 12JAN93 B 3 3 Polychaeta 18 1.22 12JAN93 B 3 10 Rhynchocoela 1 0.03 12JAN93 B 3 10 Polychaeta 10 1.56 12JAN93 C 1 3 Crustacea 2 0.04 12JAN93 C 1 3 Mollusca 5 1.52 12JAN93 C 1 3 Polychaeta 44 2.99 12JAN93 C 1 10 Rhynchocoela 2 0.28 12JAN93 C 1 10 Polychaeta 6 0.98 12JAN93 C 2 3 Crustacea 4 0.06 12JAN93 C 2 3 Mollusca 1 0.03 12JAN93 C 2 3 Polychaeta 25 4.04 12JAN93 C 2 10 Polychaeta 5 0.73 12JAN93 C 3 3 Crustacea 3 0.09 12JAN93 C 3 3 Mollusca 4 0.4 12JAN93 C 3 3 Rhynchocoela 1 0.06 12JAN93 C 3 3 Polychaeta 30 2.12 12JAN93 C 3 10 Polychaeta 1 0.14 12JAN93 D 1 3 Mollusca 15 0.95 12JAN93 D 1 3 Polychaeta 37 4.09 12JAN93 D 1 10 Polychaeta 1 0.1 12JAN93 D 2 3 Mollusca 42 5.02 12JAN93 D 2 3 Other 1 3.8 12JAN93 D 2 3 Polychaeta 31 2.45 12JAN93 D 2 10 Mollusca 2 0.06 12JAN93 D 2 10 Rhynchocoela 1 0.02 12JAN93 D 2 10 Polychaeta 2 0.13 12JAN93 D 3 3 Mollusca 68 11.77 12JAN93 D 3 3 Rhynchocoela 3 0.3 12JAN93 D 3 3 Polychaeta 17 1.02 12JAN93 D 3 10 Polychaeta 3 0.24 05APR93 A 1 3 Chironomid larvae 1 0.58 05APR93 A 1 3 Mollusca 162 16.81 05APR93 A 1 3 Polychaeta 11 1.82 05APR93 A 1 10 Mollusca 3 0.35 05APR93 A 1 10 Polychaeta 2 0.42 05APR93 A 2 3 Mollusca 180 24.78 05APR93 A 2 3 Rhynchocoela 1 0.47 05APR93 A 2 3 Polychaeta 20 2.5 05APR93 A 2 10 Mollusca 1 0.29 05APR93 A 2 10 Polychaeta 3 0.82 05APR93 A 3 3 Mollusca 120 13.43 05APR93 A 3 3 Polychaeta 8 2.23 05APR93 A 3 10 Mollusca 5 0.56 05APR93 A 3 10 Rhynchocoela 1 0.47 05APR93 A 3 10 Polychaeta 3 0.39 05APR93 B 1 3 Mollusca 110 12.91 05APR93 B 1 3 Polychaeta 119 12.15 05APR93 B 1 10 Mollusca 1 6.88 05APR93 B 1 10 Polychaeta 14 4.34 05APR93 B 2 3 Mollusca 41 3.52 05APR93 B 2 3 Polychaeta 141 11.5 05APR93 B 2 10 Mollusca 3 0.24 05APR93 B 2 10 Polychaeta 16 3.08 05APR93 B 3 3 Mollusca 100 15.62 05APR93 B 3 3 Polychaeta 91 9.27 05APR93 B 3 10 Mollusca 3 20.71 05APR93 B 3 10 Rhynchocoela 1 0.39 05APR93 B 3 10 Polychaeta 14 2.19 05APR93 C 1 3 Mollusca 95 20.36 05APR93 C 1 3 Polychaeta 68 4.53 05APR93 C 1 10 Polychaeta 4 0.56 05APR93 C 2 3 Mollusca 160 34.91 05APR93 C 2 3 Polychaeta 98 5.3 05APR93 C 2 10 Polychaeta 1 0.27 05APR93 C 3 3 Mollusca 43 9.16 05APR93 C 3 3 Rhynchocoela 1 0.51 05APR93 C 3 3 Polychaeta 63 3.9 05APR93 C 3 10 Rhynchocoela 2 3.41 05APR93 C 3 10 Polychaeta 15 3.17 05APR93 D 1 3 Mollusca 39 17.21 05APR93 D 1 3 Polychaeta 36 2.8 05APR93 D 1 10 Mollusca 1 0.22 05APR93 D 1 10 Rhynchocoela 1 0.14 05APR93 D 1 10 Polychaeta 11 1.1 05APR93 D 2 3 Crustacea 1 0.1 05APR93 D 2 3 Mollusca 31 16.96 05APR93 D 2 3 Rhynchocoela 1 0.06 05APR93 D 2 3 Polychaeta 36 1.7 05APR93 D 2 10 Mollusca 3 0.06 05APR93 D 2 10 Other 1 1.29 05APR93 D 2 10 Polychaeta 1 0.95 05APR93 D 3 3 Crustacea 3 0.03 05APR93 D 3 3 Mollusca 25 12.97 05APR93 D 3 3 Polychaeta 58 2.69 05APR93 D 3 10 Polychaeta 6 0.6 09JUL93 A 1 3 Chironomid larvae 3 0.1 09JUL93 A 1 3 Mollusca 49 6.94 09JUL93 A 1 3 Polychaeta 7 0.16 09JUL93 A 1 10 Polychaeta 3 0.11 09JUL93 A 2 3 Mollusca 100 11.56 09JUL93 A 2 3 Polychaeta 4 0.08 09JUL93 A 2 10 Mollusca 1 0.05 09JUL93 A 3 3 Mollusca 91 13.31 09JUL93 A 3 3 Polychaeta 8 0.07 09JUL93 A 3 10 Polychaeta 1 0.01 09JUL93 B 1 3 Mollusca 70 7.24 09JUL93 B 1 3 Polychaeta 2 0.02 09JUL93 B 1 10 Mollusca 5 0.59 09JUL93 B 1 10 Polychaeta 9 1.1 09JUL93 B 2 3 Mollusca 39 5.49 09JUL93 B 2 3 Polychaeta 5 0.25 09JUL93 B 2 10 Mollusca 7 0.82 09JUL93 B 2 10 Polychaeta 22 3.42 09JUL93 B 3 3 Mollusca 55 6.87 09JUL93 B 3 3 Polychaeta 8 2.77 09JUL93 B 3 10 Mollusca 1 0.07 09JUL93 B 3 10 Polychaeta 21 1.83 09JUL93 C 1 3 Mollusca 2 0.23 09JUL93 C 1 3 Polychaeta 4 3.42 09JUL93 C 1 10 Polychaeta 17 5.22 09JUL93 C 2 3 Crustacea 1 0.08 09JUL93 C 2 3 Mollusca 13 4.44 09JUL93 C 2 3 Polychaeta 11 3.6 09JUL93 C 2 10 Mollusca 3 1.25 09JUL93 C 2 10 Polychaeta 33 6.47 09JUL93 C 3 3 Mollusca 26 6.54 09JUL93 C 3 3 Polychaeta 16 2.28 09JUL93 C 3 10 Rhynchocoela 1 1.3 09JUL93 C 3 10 Polychaeta 16 2.45 09JUL93 D 1 3 Mollusca 3 0.42 09JUL93 D 1 3 Polychaeta 5 0.43 09JUL93 D 1 10 Polychaeta 10 2.41 09JUL93 D 2 3 Polychaeta 8 0.28 09JUL93 D 2 10 Polychaeta 0 0 09JUL93 D 3 3 Polychaeta 17 0.89 09JUL93 D 3 10 Polychaeta 3 0.56 110CT93 A 1 3 Mollusca 25 16.99 110CT93 A 1 3 Polychaeta 15 0.69 110CT93 A 1 10 Polychaeta 0 0 110CT93 A 2 3 Mollusca 21 14.82 110CT93 A 2 3 Polychaeta 11 0.63 110CT93 A 2 10 Polychaeta 2 0.36 110CT93 A 3 3 Mollusca 20 9.86 110CT93 A 3 3 Other 1 0.02 110CT93 A 3 3 Polychaeta 9 0.96 110CT93 A 3 10 Polychaeta 1 0.2 110CT93 B 1 3 Mollusca 2 2.26 110CT93 B 1 3 Polychaeta 3 0.81 110CT93 B 1 10 Polychaeta 8 2.6 110CT93 B 2 3 Mollusca 7 6.66 110CT93 B 2 3 Polychaeta 7 1.59 110CT93 B 2 10 Polychaeta 6 1.17 110CT93 B 3 3 Mollusca 17 15.59 110CT93 B 3 3 Rhynchocoela 1 0.47 110CT93 B 3 3 Other 1 0.03 110CT93 B 3 3 Polychaeta 11 0.81 110CT93 B 3 10 Polychaeta 16 2.99 110CT93 C 1 3 Crustacea 15 4 110CT93 C 1 3 Mollusca 63 152.97 110CT93 C 1 3 Rhynchocoela 1 0.22 110CT93 C 1 3 Polychaeta 15 1.07 110CT93 C 1 10 Polychaeta 6 0.59 110CT93 C 2 3 Mollusca 14 3.82 110CT93 C 2 3 Other 1 0.2 110CT93 C 2 3 Polychaeta 11 0.92 110CT93 C 2 10 Polychaeta 22 3.56 110CT93 C 3 3 Mollusca 6 9.02 110CT93 C 3 3 Polychaeta 6 0.53 110CT93 C 3 10 Polychaeta 3 0.11 110CT93 D 1 3 Crustacea 1 0.01 110CT93 D 1 3 Rhynchocoela 2 0.24 110CT93 D 1 3 Polychaeta 8 0.94 110CT93 D 1 10 Polychaeta 6 0.9 110CT93 D 2 3 Mollusca 4 0.61 110CT93 D 2 3 Rhynchocoela 1 0.05 110CT93 D 2 3 Polychaeta 8 0.41 110CT93 D 2 10 Mollusca 1 0.18 110CT93 D 2 10 Polychaeta 2 0.26 110CT93 D 3 3 Rhynchocoela 1 0.05 110CT93 D 3 3 Other 1 0.11 110CT93 D 3 3 Polychaeta 7 0.95 110CT93 D 3 10 Mollusca 1 0.12 110CT93 D 3 10 Polychaeta 1 0.04 05JAN94 A 1 3 Mollusca 19 4.5 05JAN94 A 1 3 Other 4 0.02 05JAN94 A 1 3 Polychaeta 8 2.53 05JAN94 A 1 10 Polychaeta 1 0.58 05JAN94 A 2 3 Mollusca 12 7.96 05JAN94 A 2 3 Polychaeta 14 1.81 05JAN94 A 2 10 Polychaeta 5 8.61 05JAN94 A 3 3 Chironomid larvae 1 0.14 05JAN94 A 3 3 Mollusca 11 20.37 05JAN94 A 3 3 Other 1 0.01 05JAN94 A 3 3 Polychaeta 6 0.53 05JAN94 A 3 10 Chironomid larvae 1 0.37 05JAN94 A 3 10 Polychaeta 2 0.46 05JAN94 B 1 3 Polychaeta 15 1.81 05JAN94 B 1 10 Polychaeta 4 14.54 05JAN94 B 2 3 Mollusca 3 7.42 05JAN94 B 2 3 Other 1 0.01 05JAN94 B 2 3 Polychaeta 16 1.57 05JAN94 B 2 10 Polychaeta 2 1.4 05JAN94 B 3 3 Chironomid larvae 1 0.18 05JAN94 B 3 3 Polychaeta 19 1.41 05JAN94 B 3 10 Polychaeta 6 1.72 05JAN94 C 1 3 Mollusca 7 4.22 05JAN94 C 1 3 Rhynchocoela 3 1.2 05JAN94 C 1 3 Polychaeta 19 1.81 05JAN94 C 1 10 Polychaeta 3 0.74 05JAN94 C 2 3 Crustacea 2 0.06 05JAN94 C 2 3 Mollusca 13 8.16 05JAN94 C 2 3 Rhynchocoela 2 0.07 05JAN94 C 2 3 Other 1 0.01 05JAN94 C 2 3 Polychaeta 34 2.34 05JAN94 C 2 10 Mollusca 1 0.2 05JAN94 C 2 10 Polychaeta 1 0.25 05JAN94 C 3 3 Mollusca 13 8 05JAN94 C 3 3 Polychaeta 43 2.72 05JAN94 C 3 10 Polychaeta 1 0.14 05JAN94 D 1 3 Mollusca 6 0.84 05JAN94 D 1 3 Rhynchocoela 1 0.29 05JAN94 D 1 3 Polychaeta 27 2.92 05JAN94 D 1 10 Polychaeta 2 0.24 05JAN94 D 2 3 Crustacea 1 0.26 05JAN94 D 2 3 Mollusca 1 0.23 05JAN94 D 2 3 Rhynchocoela 2 0.22 05JAN94 D 2 3 Other 1 0.24 05JAN94 D 2 3 Polychaeta 26 2.79 05JAN94 D 2 10 Polychaeta 4 0.97 05JAN94 D 3 3 Other 1 0.09 05JAN94 D 3 3 Polychaeta 26 2.96 05JAN94 D 3 10 Polychaeta 4 2.97 07APR94 A 1 3 Crustacea 3 0.09 07APR94 A 1 3 Mollusca 19 5.07 07APR94 A 1 3 Polychaeta 99 8.16 07APR94 A 1 10 Polychaeta 6 3.85 07APR94 A 2 3 Crustacea 3 0.21 07APR94 A 2 3 Mollusca 25 3.87 07APR94 A 2 3 Polychaeta 59 4.73 07APR94 A 2 10 Polychaeta 2 2.4 07APR94 A 3 3 Chironomid larvae 1 0.29 07APR94 A 3 3 Mollusca 22 4.2 07APR94 A 3 3 Polychaeta 44 4.71 07APR94 A 3 10 Polychaeta 2 3.98 07APR94 B 1 3 Mollusca 10 0.39 07APR94 B 1 3 Polychaeta 99 5.57 07APR94 B 1 10 Polychaeta 9 3.4 07APR94 B 2 3 Mollusca 4 0.25 07APR94 B 2 3 Polychaeta 138 8.76 07APR94 B 2 10 Polychaeta 15 5.63 07APR94 B 3 3 Crustacea 1 0.05 07APR94 B 3 3 Mollusca 4 0.12 07APR94 B 3 3 Rhynchocoela 1 0.01 07APR94 B 3 3 Other 1 0.1 07APR94 B 3 3 Polychaeta 106 7.67 07APR94 B 3 10 Polychaeta 3 2.01 07APR94 C 1 3 Crustacea 3 0.04 07APR94 C 1 3 Mollusca 14 12.82 07APR94 C 1 3 Polychaeta 51 4.96 07APR94 C 1 10 Polychaeta 9 2.12 07APR94 C 2 3 Crustacea 3 0.15 07APR94 C 2 3 Mollusca 11 5.75 07APR94 C 2 3 Polychaeta 49 5.82 07APR94 C 2 10 Polychaeta 16 3.27 07APR94 C 3 3 Crustacea 3 0.05 07APR94 C 3 3 Mollusca 13 2.39 07APR94 C 3 3 Polychaeta 42 2.67 07APR94 C 3 10 Mollusca 1 14.25 07APR94 C 3 10 Polychaeta 2 1.31 07APR94 D 1 3 Crustacea 2 0.05 07APR94 D 1 3 Mollusca 1 0.2 07APR94 D 1 3 Polychaeta 18 1.23 07APR94 D 1 10 Polychaeta 8 5.06 07APR94 D 2 3 Polychaeta 22 3.61 07APR94 D 2 10 Polychaeta 6 6.61 07APR94 D 3 3 Rhynchocoela 1 0.14 07APR94 D 3 3 Other 1 0.02 07APR94 D 3 3 Polychaeta 50 3.42 07APR94 D 3 10 Polychaeta 6 0.28 SPECIES DATA Macrofauna Number species found in biological samples. per core, multiply by 183 to obtain number per m 2. Lavaca-Colorado Estuary Date STA REP SEC Species n 060CT92 A 1 3 Pseudodiaptomus coronatus 1 060CT92 A 1 3 Mediomastus ambiseta 14 060CT92 A 1 10 Rhynchocoel (unidentified) 1 060CT92 A 1 10 Mediomastus ambiseta 10 060CT92 A 2 3 Streblospio benedicti 3 060CT92 A 2 3 Mediomastus ambiseta 14 060CT92 A 2 10 No species observed 0 060CT92 A 3 3 Streblospio benedicti 2 060CT92 A 3 3 Mediomastus ambiseta 26 060CT92 A 3 10 Rhynchocoel (unidentified) 1 060CT92 A 3 10 Oligochaetes (unidentified) 1 060CT92 A 3 10 Streblospio benedicti 1 060CT92 A 3 10 Chironomid larvae 1 060CT92 A 3 10 Parandalia ocularis 1 060CT92 A 3 10 Mediomastus ambiseta 9 060CT92 B 1 3 Streblospio benedicti 3 060CT92 B 1 3 Mulinia lateralis 1 060CT92 B 1 3 Mediomastus ambiseta 16 060CT92 B 1 10 Mediomastus ambiseta 6 060CT92 B 2 3 Streblospio benedicti 3 060CT92 B 2 3 Mediomastus ambiseta 16 060CT92 B 2 10 Glycinde solitaria 1 060CT92 B 2 10 Mediomastus ambiseta 3 060CT92 B 3 3 Streblospio benedicti 1 060CT92 B 3 3 Mediomastus ambiseta 2 060CT92 B 3 10 Paraprionospio pinnata 1 060CT92 B 3 10 Mediomastus ambiseta 1 060CT92C 1 3 1 Rhynchocoel (unidentified) 060CT92 C 1 3 Anaitides erythrophyllus 1 060CT92 C 1 3 Streblospio benedicti 4 060CT92 C 1 3 Paraprionospio pinnata 1 060CT92 C 1 3 Pectinaria gouldii 1 060CT92 C 1 3 Mulinia lateralis 28 060CT92 C 13 Ophiuroidea (unidentified) 2 060CT92 C 1 3 Macoma mitchelli 1 060CT92 C 1 3 Pyramidella sp. 1 060CT92 C 1 3 Mediomastus ambiseta 44 060CT92 C 1 10 Rhynchocoel (unidentified) 1 060CT92 C 1 10 Oligochaetes (unidentified) 1 060CT92 C 1 10 Glycinde solitaria 1 060CT92 C 1 10 Paraprionospio pinnata 2 060CT92 C 1 10 Haploscoloplos foliosus 2 060CT92 C 1 10 Pinnotheridae (unidentified) 1 060CT92 C 1 10 Mediomastus ambiseta 18 060CT92 C 2 3 Oligochaetes (unidentified) 1 060CT92 C 2 3 Streblospio benedicti 2 060CT92 C 2 3 Paraprionospio pinnata 2 060CT92 C 2 3 Cossura delta 1 060CT92 C 2 3 Clymenella torquata 1 060CT92 C 2 3 Nuculana acuta 1 060CT92 C 2 3 Mulinia lateralis 16 060CT92 C 2 3 Ancistrosyllis groenlandica 1 060CT92 C 2 3 Mediomastus ambiseta 29 060CT92 C 2 10 Rhynchocoel (unidentified) 2 060CT92 C 2 10 Streblospio benedicti 1 060CT92 C 2 10 Ancistrosyllis groenlandica 1 060CT92 C 2 10 Mediomastus ambiseta 11 060CT92 C 3 3 Rhynchocoel (unidentified) 2 060CT92 C 3 3 Streblospio benedicti 4 060CT92 C 3 3 Paraprionospio pinnata 1 060CT92 C 3 3 Pectinaria gouldii 1 060CT92 C 3 3 Mulinia lateralis 22 060CT92 C 3 3 Mediomastus ambiseta 21 060CT92 C 3 10 Rhynchocoel (unidentified) 2 060CT92 C 3 10 Gyptis vittata 2 060CT92 C 3 10 Paraprionospio pinnata 2 060CT92 C 3 10 Mediomastus ambiseta 7 060CT92 D 1 3 Oligochaetes (unidentified) 1 060CT92 D 1 3 Sigambra tentaculata 1 060CT92 D 1 3 Streblospio benedicti 3 060CT92 D 1 3 Minuspio cirrifera 3 060CT92 D 1 3 Cossura delta 11 060CT92 D 1 3 Nuculana acuta 1 060CT92 D 1 3 Ophiuroidea (unidentified) 1 060CT92 D 1 3 Mediomastus ambiseta 19 060CT92 D 1 10 Rhynchocoel (unidentified) 3 060CT92 D 1 10 Oligochaetes (unidentified) 1 060CT92 D 1 10 Gyptis vittata 1 060CT92 D 1 10 Minuspio cirrifera 2 060CT92 D 1 10 Corbula contracta 1 060CT92 D 1 10 Paraonidae Grp. B 1 060CT92 D 1 10 Ophiuroidea (unidentified) 1 060CT92 D 1 10 Apseudes sp. A 3 060CT92 D 1 10 Naineris sp. A 1 060CT92 D 1 10 Paramya subovata 4 060CT92 D 2 3 Drilonereis magna 2 060CT92 D 2 3 Streblospio benedicti 2 060CT92 D 2 3 Paraprionospio pinnata 1 060CT92 D 2 3 Cossura delta 10 060CT92 D 2 3 Melinna maculata 1 060CT92 D 2 3 Nuculana concentrica 1 060CT92 D 2 3 Mediomastus ambiseta 42 060CT92 D 2 3 Paramya subovata 1 060CT92 D 2 10 Rhynchocoel (unidentified) 1 060CT92 D 2 10 Oligochaetes (unidentified) 1 060CT92 D 2 10 Paraprionospio pinnata 1 060CT92 D 2 10 Minuspio cirrifera 1 060CT92 D 2 10 Cossura delta 5 060CT92 D 2 10 Paraonidae Grp. B 1 060CT92 D 2 10 Ophiuroidea (unidentified) 1 060CT92 D 2 10 Mediomastus ambiseta 6 060CT92 D 2 10 Paramya subovata 1 060CT92 D 3 3 Streblospio benedicti 5 060CT92 D 3 3 Paraprionospio pinnata 1 060CT92 D 3 3 Minuspio cirrifera 1 060CT92 D 3 3 Cossura delta 2 060CT92 D 3 3 Nuculana concentrica 1 060CT92 D 3 3 Mediomastus ambiseta 29 060CT92 D 3 10 Rhynchocoel (unidentified) 2 060CT92 D 3 10 Sigambra tentaculata 1 060CT92 D 3 10 Minuspio cirrifera 3 060CT92 D 3 10 Ophiuroidea (unidentified) 2 060CT92 D 3 10 Periploma cf. orbiculare 1 060CT92 D 3 10 Mediomastus ambiseta 8 060CT92 D 3 10 Paramya subovata 7 12JAN93 A 1 3 Streblospio benedicti 7 12JAN93 A 1 3 Cossura delta 1 12JAN93 A 1 3 Capitella capitata 1 12JAN93 A 1 3 Mulinia lateralis 1 12JAN93 A 1 3 Chironomid larvae 1 12JAN93 A 1 3 Macoma mitchelli 3 12JAN93 A 1 3 Mediomastus ambiseta 33 12JAN93 A 1 10 Glycinde solitaria 1 12JAN93 A 2 3 Diopatra cuprea 1 12JAN93 A 2 3 Streblospio benedicti 4 12JAN93 A 2 3 Cossura delta 1 12JAN93 A 2 3 Capitella capitata 3 12JAN93 A 2 3 Mulinia lateralis 1 12JAN93 A 2 3 Macoma mitchelli 2 12JAN93 A 2 3 Mediomastus ambiseta 73 12JAN93 A 2 10 Glycinde solitaria 1 12JAN93 A 2 10 Mulinia lateralis 1 12JAN93 A 2 10 Laeonereis culveri 1 12JAN93 A 2 10 Mediomastus ambiseta 1 12JAN93 A 3 3 Streblospio benedicti 6 12JAN93 A 3 3 Mulinia lateralis 1 12JAN93 A 3 3 Macoma mitchelli 2 12JAN93 A 3 3 Mediomastus ambiseta 26 12JAN93 A 3 10 No species observed 0 12JAN93 B 1 3 Streblospio benedicti 4 12JAN93 B 1 3 Haploscoloplos foliosus 1 12JAN93 B 1 3 Cossura delta 1 12JAN93 B 1 3 Mulinia lateralis 2 12JAN93 B 1 3 Pyramidella sp. 1 12JAN93 B 1 3 Mediomastus ambiseta 11 12JAN93 B 1 10 Mediomastus ambiseta 2 12JAN93 B 2 3 Rhynchocoel (unidentified) 1 12JAN93 B 2 3 Glycinde solitaria 1 12JAN93 B 2 3 Streblospio benedicti 8 12JAN93 B 2 3 Cossura delta 2 12JAN93 B 2 3 Macoma mitchelli 3 12JAN93 B 2 3 Mediomastus ambiseta 28 12JAN93 B 2 10 Mediomastus ambiseta 4 12JAN93 B 3 3 Streblospio benedicti 5 12JAN93 B 3 3 Haploscoloplos foliosus 1 12JAN93 B 3 3 Cossura delta 4 12JAN93 B 3 3 Monoculodes sp. 1 12JAN93 B 3 3 Macoma mitchelli 1 12JAN93 B 3 3 Mediomastus ambiseta 23 12JAN93 B 3 10 Cossura delta 1 12JAN93 B 3 10 Mediomastus ambiseta 1 12JAN93 C 1 3 Rhynchocoel (unidentified) 1 12JAN93 C 1 3 Glycinde solitaria 6 12JAN93 C 1 3 Streblospio benedicti 2 12JAN93 C 1 3 Haploscoloplos foliosus 1 12JAN93 C 1 3 Cossura delta 3 12JAN93 C 1 3 Axiothella mucosa 1 12JAN93 C 1 3 Mulinia lateralis 1 12JAN93 C 1 3 Lyonsia hyalina floridana 1 12JAN93 C 1 3 Pseudodiaptomus coronatus 1 12JAN93 C 1 3 Cyclaspis varians 1 12JAN93 C 1 3 Nassarius acutus 1 12JAN93 C 1 3 Sigalionidae (unidentified) 1 12JAN93 C 1 3 Ophiuroidea (unidentified) 3 12JAN93 C 1 3 Mediomastus ambiseta 71 12JAN93 C 1 10 Glycinde solitaria 1 12JAN93 C 1 10 Haploscoloplos foliosus 4 12JAN93 C 1 10 Cossura delta 2 12JAN93 C 1 10 Turbellaria (unidentified) 1 12JAN93 C 1 10 Mediomastus ambiseta 11 12JAN93 C 2 3 Rhynchocoel (unidentified) 1 12JAN93 C 2 3 Oligochaetes (unidentified) 1 12JAN93 C 2 3 Eteone heteropoda 1 12JAN93 C 2 3 Gyptis vittata 1 12JAN93 C 2 3 Glycinde solitaria 6 12JAN93 C 2 3 Haploscoloplos foliosus 2 12JAN93 C 2 3 Cossura delta 1 12JAN93 C 2 3 Axiothella mucosa 1 12JAN93 C 2 3 Pectinaria gouldii 1 12JAN93 C 2 3 Cyclaspis varians 1 12JAN93 C 2 3 Oxyurostylis sp. 1 12JAN93 C 2 3 Mediomastus ambiseta 37 12JAN93 C 2 10 Glycinde solitaria 1 12JAN93 C 2 10 Haploscoloplos foliosus 2 12JAN93 C 2 10 Cossura delta 1 12JAN93 C 2 10 Paraonidae Grp. A 1 12JAN93 C 2 10 Mediomastus ambiseta 3 12JAN93 C 3 3 Rhynchocoel (unidentified) 2 12JAN93 C 3 3 Gyptis vittata 1 12JAN93 C 3 3 Glycinde solitaria 3 12JAN93 C 3 3 Streblospio benedicti 2 12JAN93 C 3 3 Cossura delta 3 12JAN93 C 3 3 Pectinaria gouldii 1 12JAN93 C 3 3 Mulinia lateralis 1 12JAN93 C 3 3 Phascolion strombi 1 12JAN93 C 3 3 Ophiuroidea (unidentified) 3 12JAN93 C 3 3 Mediomastus ambiseta 39 12JAN93 C 3 10 Rhynchocoel (unidentified) 2 12JAN93 C 3 10 Cossura delta 4 12JAN93 C 3 10 Mediomastus ambiseta 8 12JAN93 D 1 3 Rhynchocoel (unidentified) 3 12JAN93 D 1 3 Eunoe cf. nodulosa 1 12JAN93 D 1 3 Sthenelais boa 1 12JAN93 D 1 3 Glycinde solitaria 2 12JAN93 D 13 Streblospio benedicti 1 12JAN93 D 1 3 Minuspio cirrifera 4 12JAN93 D 1 3 Cossura delta 3 12JAN93 D 1 3 Pectinaria gouldii 1 12JAN93 D 1 3 Melinna maculata 1 12JAN93 D 1 3 Corbula contracta 1 12JAN93 D 1 3 Nuculana concentrica 1 12JAN93 D 1 3 Ophiuroidea (unidentified) 11 12JAN93 D 1 3 Pelecypoda (unidentified) 8 12JAN93 D 1 3 Mediomastus ambiseta 41 12JAN93 D 1 3 Munna sp. 1 12JAN93 D 1 10 Sigambra tentaculata 1 12JAN93 D 1 10 Ophiuroidea (unidentified) 1 12JAN93 D 1 10 Mediomastus ambiseta 3 12JAN93 D 2 3 Rhynchocoel (unidentified) 1 12JAN93 D 2 3 Sthenelais boa 1 12JAN93 D 2 3 Sigambra tentaculata 1 12JAN93 D 2 3 Glycinde solitaria 2 12JAN93 D 2 3 Mediomastus ambiseta 9 12JAN93 D 2 10 Naineris sp. A 3 12JAN93 D 2 10 Mediomastus ambiseta 2 12JAN93 D 3 3 Anthozoa (unidentified) 1 12JAN93 D 3 3 Rhynchocoel (unidentified) 2 12JAN93 D 3 3 Oligochaetes (unidentified) 1 12JAN93 D 3 3 Glycinde solitaria 2 12JAN93 D 3 3 Drilonereis magna 1 12JAN93 D 3 3 Cossura delta 5 12JAN93 D 3 3 Ophiuroidea (unidentified) 1 12JAN93 D 3 3 Apseudes sp. A 2 12JAN93 D 3 3 Mediomastus ambiseta 14 12JAN93 D 3 10 Clymenella torquata 1 12JAN93 D 3 10 Owenia fusiformis 1 12JAN93 D 3 10 Sigambra cf. wassi 1 12JAN93 D 3 10 Mediomastus ambiseta 3 12JAN93 E 1 3 Rhynchocoel (unidentified) 2 12JAN93 E 1 3 Oligochaetes (unidentified) 1 12JAN93 E 1 3 Gyptis vittata 1 12JAN93 E 1 3 Streblospio benedicti 10 12JAN93 E 1 3 Paraprionospio pinnata 2 12JAN93 E 1 3 Nuculana concentrica 1 12JAN93 E 1 3 Ophiuroidea (unidentified) 1 12JAN93 E 1 3 Pyramidella crenulata 1 12JAN93 E 1 3 Mediomastus ambiseta 47 12JAN93 E 1 10 Rhynchocoel (unidentified) 1 12JAN93 E 1 10 Gyptis vittata 1 12JAN93 E 1 10 Streblospio benedicti 1 12JAN93 E 1 10 Paraprionospio pinnata 2 12JAN93 E 1 10 Cossura delta 2 12JAN93 E 1 10 Ophiuroidea (unidentified) 1 12JAN93 E 1 10 Mediomastus ambiseta 2 12JAN93 E 2 3 Oligochaetes (unidentified) 1 12JAN93 E 2 3 Streblospio benedicti 7 12JAN93 E 2 3 Cossura delta 2 12JAN93 E 2 3 Asychis sp. 1 12JAN93 E 2 3 Pyramidella crenulata 2 12JAN93 E 2 3 Mediomastus ambiseta 31 12JAN93 E 2 10 Gyptis vittata 2 12JAN93 E 2 10 Glycinde solitaria 1 12JAN93 E 2 10 Paraprionospio pinnata 8 12JAN93 E 2 10 Paraonidae Grp. A 1 12JAN93 E 3 3 Glycinde solitaria 3 12JAN93 E 3 3 Streblospio benedicti 4 12JAN93 E 3 3 Paraprionospio pinnata 4 12JAN93 E 3 3 Paraonidae Grp. B 1 12JAN93 E 3 3 Ophiuroidea (unidentified) 1 12JAN93 E 3 3 Mediomastus ambiseta 19 12JAN93 E 3 10 Glycinde solitaria 3 12JAN93 E 3 10 Streblospio benedicti 1 12JAN93 E 3 10 Paraprionospio pinnata 1 12JAN93 E 3 10 Paraonidae Grp. A 1 12JAN93 E 3 10 Mediomastus ambiseta 1 12JAN93 F 1 3 Streblospio benedicti 1 12JAN93 F 1 3 Mulinia lateralis 2 12JAN93 F 1 3 Macoma mitchelli 2 12JAN93 F 1 3 Mediomastus ambiseta 25 12JAN93 F 1 10 Paraprionospio pinnata 1 12JAN93 F 1 10 Cossura delta 1 12JAN93 F 1 10 Mediomastus ambiseta 1 12JAN93 F 2 3 Anthozoa (unidentified) 1 12JAN93 F 2 3 Rhynchocoel (unidentified) 1 12JAN93 F 2 3 Streblospio benedicti 3 12JAN93 F 2 3 Haploscoloplos foliosus 2 12JAN93 F 2 3 Mulinia lateralis 6 12JAN93 F 2 3 Macoma mitchelli 3 12JAN93 F 2 3 Turbellaria (unidentified) 1 12JAN93 F 2 3 Mediomastus ambiseta 55 12JAN93 F 2 10 Rhynchocoel (unidentified) 2 12JAN93 F 2 10 Gyptis vittata 1 12JAN93 F 2 10 Paraprionospio pinnata 1 12JAN93 F 2 10 Mediomastus ambiseta 1 12JAN93 F 3 3 Streblospio benedicti 7 12JAN93 F 3 3 Paraprionospio pinnata 1 12JAN93 F 3 3 Mulinia lateralis 4 12JAN93 F 3 3 Macoma mitchelli 2 12JAN93 F 3 3 Mediomastus ambiseta 30 12JAN93 F 3 10 Oligochaetes (unidentified) 1 12JAN93 F 3 10 Paraprionospio pinnata 1 12JAN93 F 3 10 Mediomastus ambiseta 2 05APR93 A 1 3 Anthozoa (unidentified) 1 05APR93 A 1 3 Rhynchocoel (unidentified) 1 05APR93 A 1 3 Streblospio benedicti 4 05APR93 A 1 3 Mulinia lateralis 1 05APR93 A 1 3 Mediomastus ambiseta 44 05APR93 A 1 10 Macoma mitchelli 1 05APR93 A 1 10 Mediomastus ambiseta 4 05APR93 A 2 3 Streblospio benedicti 4 05APR93 A 2 3 Mediomastus ambiseta 9 05APR93 A 2 10 Macoma mitchelli 1 05APR93 A 2 10 Mediomastus ambiseta 1 05APR93 A 3 3 Rhynchocoel (unidentified) 5 05APR93 A 3 3 Streblospio benedicti 4 05APR93 A 3 3 Hobsonia florida 1 05APR93 A 3 3 Mediomastus ambiseta 38 05APR93 A 3 10 Macoma mitchelli 1 05APR93 A 3 10 Mediomastus ambiseta 10 05APR93 B 1 3 Streblospio benedicti 7 05APR93 B 1 3 Cossura delta 2 05APR93 B 1 3 Capitella capitata 1 05APR93 B 1 3 Mulinia lateralis 6 05APR93 B 1 3 Brachidontes exustus 1 05APR93 B 1 3 Mediomastus ambiseta 50 05APR93 B 1 10 Rhynchocoel (unidentified) 1 05APR93 B 1 10 Glycinde solitaria 1 05APR93 B 1 10 Cossura delta 8 05APR93 B 1 10 Macoma mitchelli 1 05APR93 B 1 10 Mediomastus ambiseta 7 05APR93 B 2 3 Rhynchocoel (unidentified) 1 05APR93 B 2 3 Sigambra bassi 1 05APR93 B 2 3 Streblospio benedicti 2 05APR93 B 2 3 Mulinia lateralis 2 05APR93 B 2 3 Mediomastus ambiseta 38 05APR93 B 2 10 Rhynchocoel (unidentified) 1 05APR93 B 2 10 Cossura delta 1 05APR93 B 2 10 Turbellaria (unidentified) 4 05APR93 B 2 10 Mediomastus ambiseta 7 05APR93 B 3 3 Cossura delta 1 05APR93 B 3 3 Mediomastus ambiseta 10 05APR93 B 3 10 Rhynchocoel (unidentified) 2 05APR93 B 3 10 Paraprionospio pinnata 1 05APR93 B 3 10 Cossura delta 4 05APR93 B 3 10 Mediomastus ambiseta 4 05APR93 C 1 3 Glycinde solitaria 2 05APR93 C 1 3 Mulinia lateralis 32 05APR93 C 1 3 Edotea montosa 1 05APR93 C 1 3 Nuculana concentrica 1 05APR93 C 1 3 Mediomastus ambiseta 2 05APR93 C 1 10 Glycinde solitaria 1 05APR93 C 1 10 Cossura delta 2 05APR93 C 1 10 Mulinia lateralis 1 05APR93 C 1 10 Lyonsia hyalina floridana 1 05APR93 C 1 10 Mediomastus ambiseta 1 05APR93 C 2 3 Capitella capitata 1 05APR93 C 2 3 Pectinaria gouldii 1 05APR93 C 2 3 Mulinia lateralis 11 05APR93 C 2 3 Cyclaspis varians 4 05APR93 C 2 3 Oxyurostylis sp. 1 05APR93 C 2 3 Mediomastus ambiseta 5 05APR93 C 2 10 Sigambra tentaculata 1 05APR93 C 2 10 Streblospio benedicti 1 05APR93 C 2 10 Mulinia lateralis 1 05APR93 C 2 10 Dorvilleidae 2 05APR93 C 2 10 Mediomastus ambiseta 4 05APR93 C 3 3 Glycinde solitaria 1 05APR93 C 3 3 Haploscoloplos foliosus 1 05APR93 C 3 3 Mulinia lateralis 9 05APR93 C 3 3 Cyclaspis varians 1 05APR93 C 3 3 Nuculana concentrica 1 05APR93 C 3 3 Oxyurostylis sp. 1 05APR93 C 3 3 Mediomastus ambiseta 1 05APR93 C 3 10 Sigambra bassi 1 05APR93 C 3 10 Gyptis vittata 1 05APR93 C 3 10 Mediomastus ambiseta 2 05APR93 D 1 3 Anthozoa (unidentified) 1 05APR93 D 1 3 Rhynchocoel (unidentified) 1 05APR93 D 1 3 Oligochaetes (unidentified) 4 05APR93 D 1 3 Glycinde solitaria 5 05APR93 D 1 3 Streblospio benedicti 1 05APR93 D 1 3 Minuspio cirrifera 4 05APR93 D 1 3 Cossura delta 6 05APR93 D 1 3 Branchioasychis americana 1 05APR93 D 1 3 Clymenella torquata 1 05APR93 D 1 3 Pectinaria gouldii 5 05APR93 D 1 3 Aligena texasiana 1 05APR93 D 1 3 Caprellid 1 05APR93 D 1 3 Hobsonia florida 6 05APR93 D 1 3 Mediomastus ambiseta 52 05APR93 D 1 10 Rhynchocoel (unidentified) 1 05APR93 D 1 10 Oligochaetes (unidentified) 1 05APR93 D 1 10 Sigambra tentaculata 1 05APR93 D 1 10 Diopatra cuprea 1 05APR93 D 1 10 Lumbrineris parvapedata 1 05APR93 D 1 10 Minuspio cirrifera 4 05APR93 D 1 10 Ophiuroidea (unidentified) 4 05APR93 D 1 10 Mediomastus ambiseta 4 05APR93 D 2 3 Anthozoa (unidentified) 2 05APR93 D 2 3 Rhynchocoel (unidentified) 1 05APR93 D 2 3 Minuspio cirrifera 2 05APR93 D 2 3 Cossura delta 7 05APR93 D 2 3 Phascolion strombi 1 05APR93 D 2 3 Ancistrosyllis groenlandica 1 05APR93 D 2 3 Ophiuroidea (unidentified) 1 05APR93 D 2 3 Hobsonia florida 1 05APR93 D 2 3 Mediomastus ambiseta 61 05APR93 D 2 10 Rhynchocoel (unidentified) 4 05APR93 D 2 10 Oligochaetes (unidentified) 1 05APR93 D 2 10 Gyptis vittata 1 05APR93 D 2 10 Minuspio cirrifera 2 05APR93 D 2 10 Naineris sp. A 1 05APR93 D 2 10 Mediomastus ambiseta 2 05APR93 D 3 3 Paranaitis speciosa 1 05APR93 D 3 3 Streblospio benedicti 4 05APR93 D 3 3 Minuspio cirrifera 1 05APR93 D 3 3 Cossura delta 8 05APR93 D 3 3 Pelecypoda (unidentified) 1 05APR93 D 3 3 Microprotopus spp. 1 05APR93 D 3 3 Hobsonia florida 1 05APR93 D 3 3 Mediomastus ambiseta 36 05APR93 D 3 10 Sigambra bassi 1 05APR93 D 3 10 Sigambra tentaculata 1 05APR93 D 3 10 Magelona phyllisae 1 05APR93 D 3 10 Cossura delta 3 05APR93 D 3 10 Holothuroid (unidentified) 1 05APR93 D 3 10 Mediomastus ambiseta 6 05APR93 E 1 3 Rhynchocoel (unidentified) 3 05APR93 E 1 3 Paranaitis speciosa 1 05APR93 E 1 3 Glycinde solitaria 1 05APR93 E 1 3 Streblospio benedicti 1 05APR93 E 1 3 Paraprionospio pinnata 1 05APR93 E 1 3 Mulinia lateralis 1 05APR93 E 1 3 Monoculodes sp. 1 05APR93 E 1 3 Acteocina canaliculata 1 05APR93 E 1 3 Hobsonia florida 8 05APR93 E 1 3 Mediomastus ambiseta 25 05APR93 E 1 10 Gyptis vittata 1 05APR93 E 1 10 Paraprionospio pinnata 2 05APR93 E 1 10 Paraonidae Grp. A 1 05APR93 E 1 10 Hobsonia florida 1 05APR93 E 2 3 Rhynchocoel (unidentified) 1 05APR93 E 2 3 Oligochaetes (unidentified) 1 05APR93 E 2 3 Gyptis vittata 1 05APR93 E 2 3 Diopatra cuprea 1 05APR93 E 2 3 Cossura delta 1 05APR93 E 2 3 Nuculana acuta 2 05APR93 E 2 3 Mulinia lateralis 1 05APR93 E 2 3 Acteocina canaliculata 1 05APR93 E 2 3 Nereidae (unidentified) 1 05APR93 E 2 3 Pectinariidae 1 05APR93 E 2 3 Hobsonia florida 12 05APR93 E 2 3 Mediomastus ambiseta 17 05APR93 E 2 10 Anthozoa (unidentified) 1 05APR93 E 2 10 Sigambra tentaculata 1 05APR93 E 2 10 Cossura delta 1 05APR93 E 2 10 Mediomastus ambiseta 4 05APR93 E 3 3 Oligochaetes (unidentified) 2 05APR93 E 3 3 Paranaitis speciosa 2 05APR93 E 3 3 Streblospio benedicti 2 05APR93 E 3 3 Paraprionospio pinnata 3 05APR93 E 3 3 Cossura delta 1 05APR93 E 3 3 Pectinaria gouldii 1 05APR93 E 3 3 Nuculana acuta 1 05APR93 E 3 3 Mulinia lateralis 1 05APR93 E 3 3 Ophiuroidea (unidentified) 1 05APR93 E 3 3 Pyramidella crenulata 3 05APR93 E 3 3 Hobsonia florida 13 05APR93 E 3 3 Oxyurostylis sp. 1 05APR93 E 3 3 Mediomastus ambiseta 20 05APR93 E 3 10 Rhynchocoel (unidentified) 3 05APR93 E 3 10 Lumbrineris parvapedata 1 05APR93 E 3 10 Paraprionospio pinnata 2 05APR93 E 3 10 Cossura delta 2 05APR93 E 3 10 Mediomastus ambiseta 3 05APR93 F 1 3 Streblospio benedicti 13 05APR93 F 1 3 Mulinia lateralis 9 05APR93 F 1 3 Acteocina canaliculata 1 05APR93 F 1 3 Pelecypoda (unidentified) 1 05APR93 F 1 3 Macoma mitchelli 1 05APR93 F 1 3 Turbellaria (unidentified) 4 05APR93 F 1 3 Mediomastus ambiseta 34 05APR93 F 1 10 Rhynchocoel (unidentified) 2 05APR93 F 1 10 Gyptis vittata 1 05APR93 F 1 10 Streblospio benedicti 1 05APR93 F 1 10 Paraprionospio pinnata 3 05APR93 F 1 10 Macoma mitchelli 2 05APR93 F 1 10 Turbellaria (unidentified) 2 05APR93 F 1 10 Mediomastus ambiseta 7 05APR93 F 2 3 Rhynchocoel (unidentified) 1 05APR93 F 2 3 Streblospio benedicti 8 05APR93 F 2 3 Capitella capitata 1 05APR93 F 2 3 Mulinia lateralis 5 05APR93 F 2 3 Pyramidella crenulata 1 05APR93 F 2 3 Brachidontes exustus 1 05APR93 F 2 3 Macoma mitchelli 1 05APR93 F 2 3 Mediomastus ambiseta 37 05APR93 F 2 10 Rhynchocoel (unidentified) 3 05APR93 F 2 10 Gyptis vittata 1 05APR93 F 2 10 Capitella capitata 3 05APR93 F 2 10 Mediomastus ambiseta 5 05APR93 F 3 3 Streblospio benedicti 17 05APR93 F 3 3 Capitella capitata 1 05APR93 F 3 3 Mulinia lateralis 7 05APR93 F 3 3 Mediomastus ambiseta 59 05APR93 F 3 10 Rhynchocoel (unidentified) 1 05APR93 F 3 10 Paraprionospio pinnata 1 05APR93 F 3 10 Capitella capitata 1 05APR93 F 3 10 Macoma mitchelli 2 05APR93 F 3 10 Mediomastus ambiseta 2 09JUL93 A 1 3 Streblospio benedicti 1 09JUL93 A 1 3 Mulinia lateralis 2 09JUL93 A 1 3 Littoridina sphinctostoma 1 09JUL93 A 1 3 Mediomastus ambiseta 14 09JUL93 A 1 10 Chironomid larvae 1 09JUL93 A 2 3 Streblospio benedicti 2 09JUL93 A 2 3 Mulinia lateralis 2 09JUL93 A 2 3 Chironomid larvae 1 09JUL93 A 2 3 Littoridina sphinctostoma 3 09JUL93 A 2 3 Mediomastus ambiseta 23 09JUL93 A 2 10 Macoma mitchelli 1 09JUL93 A 2 10 Mediomastus ambiseta 2 09JUL93 A 3 3 Mulinia lateralis 1 09JUL93 A 3 3 Mediomastus ambiseta 21 09JUL93 A 3 10 Capitella capitata 1 09JUL93 A 3 10 Macoma mitchelli 1 09JUL93 A 3 10 Mediomastus ambiseta 3 09JUL93 B 1 3 Streblospio benedicti 1 09JUL93 B 1 3 Chironomid larvae 1 09JUL93 B 1 3 Mediomastus ambiseta 3 09JUL93 B 1 10 Mediomastus ambiseta 2 09JUL93 B 2 3 Macoma mitchelli 1 09JUL93 B 2 3 Littoridina sphinctostoma 2 09JUL93 B 2 10 Mediomastus ambiseta 7 09JUL93 B 3 3 Streblospio benedicti 1 09JUL93 B 3 3 Mediomastus ambiseta 5 09JUL93 B 3 10 Mediomastus ambiseta 10 09JUL93 C 1 3 Streblospio benedicti 12 09JUL93 C 1 10 Capitella capitata 1 09JUL93 C 2 3 Streblospio benedicti 9 09JUL93 C 2 3 Cossura delta 1 09JUL93 C 2 3 Capitella capitata 1 09JUL93 C 2 3 Mulinia lateralis 1 09JUL93 C 2 3 Macoma mitchelli 1 09JUL93 C 2 10 Oligochaetes (unidentified) 1 09JUL93 C 2 10 Streblospio benedicti 1 09JUL93 C 2 10 Cossura delta 2 09JUL93 C 2 10 Mediomastus ambiseta 1 09JUL93 C 3 3 Streblospio benedicti 18 09JUL93 C 3 3 Mulinia lateralis 2 09JUL93 C 3 3 Parandalia ocularis 1 09JUL93 C 3 3 Mediomastus ambiseta 3 09JUL93 C 3 10 Rhynchocoel (unidentified) 1 09JUL93 C 3 10 Haploscoloplos foliosus 1 09JUL93 C 3 10 Capitella capitata 1 09JUL93 C 3 10 Axiothella mucosa 1 09JUL93 C 3 10 Ancistrosyllis groenlandica 1 09JUL93 C 3 10 Caecum johnsoni 4 09JUL93 C 3 10 Mediomastus ambiseta 1 09JUL93 D 1 3 Anthozoa (unidentified) 1 09JUL93 D 1 3 Paraprionospio pinnata 1 09JUL93 D 1 3 Minuspio cirrifera 1 09JUL93 D 1 3 Cossura delta 1 09JUL93 D 1 3 Periploma cf. orbiculare 1 09JUL93 D 1 3 Mediomastus ambiseta 4 09JUL93 D 1 3 Eudorella sp. 1 09JUL93 D 1 10 Oligochaetes (unidentified) 2 09JUL93 D 1 10 Gyptis vittata 1 09JUL93 D 1 10 Abra aequalis 1 09JUL93 D 1 10 Apseudes sp. A 2 09JUL93 D 1 10 Naineris sp. A 1 09JUL93 D 1 10 Mediomastus ambiseta 1 09JUL93 D 2 3 Diopatra cuprea 1 09JUL93 D 2 3 Paraprionospio pinnata 1 09JUL93 D 2 3 Microprotopus spp. 1 09JUL93 D 2 3 Apseudes sp. A 3 09JUL93 D 2 3 Mediomastus ambiseta 1 09JUL93 D 2 10 Rhynchocoel (unidentified) 1 09JUL93 D 2 10 Oligochaetes (unidentified) 3 09JUL93 D 2 10 Apseudes sp. A 1 09JUL93 D 2 10 Periploma cf. orbiculare 2 09JUL93 D 2 10 Naineris sp. A 1 09JUL93 D 2 10 Mediomastus ambiseta 3 09JUL93 D 3 3 Cossura delta 5 09JUL93 D 3 3 Corbula contracta 1 09JUL93 D 3 3 Mediomastus ambiseta 10 09JUL93 D 3 10 Rhynchocoel (unidentified) 1 09JUL93 D 3 10 Oligochaetes (unidentified) 1 09JUL93 D 3 10 Minuspio cirrifera 1 09JUL93 D 3 10 Cossura delta 1 09JUL93 D 3 10 Corbula contracta 1 09JUL93 D 3 10 Ophiuroidea (unidentified) 1 09JUL93 D 3 10 Periploma cf. orbiculare 2 09JUL93 D 3 10 Naineris sp. A 1 09JUL93 D 3 10 Mediomastus ambiseta 1 09JUL93 E 1 3 Glycinde solitaria 1 09JUL93 E 1 3 Streblospio benedicti 1 09JUL93 E 1 3 Paraprionospio pinnata 1 09JUL93 E 1 3 Cossura delta 2 09JUL93 E 1 3 Mulinia lateralis 10 09JUL93 E 1 3 Macoma mitchelli 1 09JUL93 E 1 3 Mediomastus ambiseta 5 09JUL93 E 1 10 Gyptis vittata 3 09JUL93 E 1 10 Paraprionospio pinnata 1 09JUL93 E 1 10 Mediomastus ambiseta 1 09JUL93 E 2 3 Glycinde solitaria 2 09JUL93 E 2 3 Streblospio benedicti 1 09JUL93 E 2 3 Cossura delta 1 09JUL93 E 2 3 Mulinia lateralis 11 09JUL93 E 2 3 Acteocina canaliculata 1 09JUL93 E 2 3 Pyramidella crenulata 2 09JUL93 E 2 3 Macoma mitchelli 1 09JUL93 E 2 3 Mediomastus ambiseta 6 09JUL93 E 2 10 Oligochaetes (unidentified) 2 09JUL93 E 2 10 Gyptis vittata 1 09JUL93 E 2 10 Paraprionospio pinnata 3 09JUL93 E 2 10 Paraonidae Grp. B 1 09JUL93 E 3 3 Rhynchocoel (unidentified) 1 09JUL93 E 3 3 Streblospio benedicti 2 09JUL93 E 3 3 Cossura delta 1 09JUL93 E 3 3 Mulinia lateralis 5 09JUL93 E 3 3 Mediomastus ambiseta 5 09JUL93 E 3 10 Oligochaetes (unidentified) 3 09JUL93 E 3 10 Gyptis vittata 1 09JUL93 E 3 10 Paraprionospio pinnata 2 09JUL93 E 3 10 Mediomastus ambiseta 1 09JUL93 F 1 3 Rhynchocoel (unidentified) 1 09JUL93 F 1 3 Gyptis vittata 1 09JUL93 F 1 3 Streblospio benedicti 2 09JUL93 F 1 3 Mediomastus ambiseta 12 09JUL93 F 1 10 Paraprionospio pinnata 3 09JUL93 F 1 10 Mediomastus ambiseta 16 09JUL93 F 2 3 Gyptis vittata 1 09JUL93 F 2 3 Paraprionospio pinnata 2 09JUL93 F 2 3 Mediomastus ambiseta 4 09JUL93 F 2 10 Glycinde solitaria 1 09JUL93 F 2 10 Paraprionospio pinnata 1 09JUL93 F 2 10 Mediomastus ambiseta 18 09JUL93 F 3 3 Streblospio benedicti 2 09JUL93 F 3 3 Paraprionospio pinnata 2 09JUL93 F 3 3 Macoma mitchelli 1 09JUL93 F 3 3 Mediomastus ambiseta 22 09JUL93 F 3 10 Glycinde solitaria 1 09JUL93 F 3 10 Mediomastus ambiseta 13 110CT93 A 1 3 Streblospio benedicti 3 110CT93 A 1 3 Mediomastus ambiseta 2 110CT93 A 1 10 No species observed 0 110CT93 A 2 3 Streblospio benedicti 6 110CT93 A 2 3 Mediomastus ambiseta 1 110CT93 A 2 10 No species observed 0 110CT93 A 3 3 No species observed 0 110CT93 A 3 10 No species observed 0 110CT93 B 1 3 Mulinia lateralis 1 110CT93 B 1 3 Mediomastus ambiseta 1 110CT93 B 1 10 Glycinde solitaria 1 11GCT93 B 1 10 Mediomastus ambiseta 2 110CT93 B 2 3 Rhynchocoel (unidentified) 2 11GCT93 B 2 3 Parandalia ocularis 2 11GCT93 B 2 3 Mediomastus ambiseta 1 110CT93 B 2 10 Mediomastus ambiseta 6 110CT93 B 3 3 Mediomastus ambiseta 1 110CT93 B 3 10 Mediomastus ambiseta 3 11GCT93 C 1 3 Rhynchocoel (unidentified) 1 110CT93 C 1 3 Streblospio benedicti 1 11GCT93 C 1 3 Paraprionospio pinnata 1 110CT93 C 1 3 Spiochaetopterus costarum 2 110CT93 C 1 3 Flaploscoloplos foliosus 1 110CT93 C 1 3 Ophiuroidea (unidentified) 1 110CT93 C 1 3 Mediomastus ambiseta 11 110CT93 C 1 10 Spiochaetopterus costarum 1 110CT93 C 1 10 Cossura delta 2 110CT93 C 1 10 Mediomastus ambiseta 9 110CT93 C 2 3 Streblospio benedicti 2 11GCT93 C 2 3 Paraprionospio pinnata 1 11GCT93 C 2 3 Spiochaetopterus costarum 1 110CT93 C 2 3 Mulinia lateralis 2 11GCT93 C 2 3 Ophiuroidea (unidentified) 2 1100T93 C 2 3 Mediomastus ambiseta 19 110CT93 C 2 10 Rhynchocoel (unidentified) 1 1100T93 C 2 10 Sigambra bassi 1 1100T93 C 2 10 Gyptis vittata 2 110CT93 C 2 10 Paraprionospio pinnata 1 110CT93 C 2 10 Haploscoloplos foliosus 1 110CT93 C 2 10 Cossura delta 1 110CT93 C 2 10 Mediomastus ambiseta 11 110CT93 C 3 3 Anthozoa (unidentified) 1 110CT93 C 3 3 Spiochaetopterus costarum 1 110CT93 C 3 3 Mulinia lateralis 2 110CT93 C 3 3 Ophiuroidea (unidentified) 1 110CT93 C 3 3 Mediomastus ambiseta 5 110CT93 C 3 10 Spiochaetopterus costarum 1 110CT93 C 3 10 Mediomastus ambiseta 1 110CT93 D 1 3 Rhynchocoel (unidentified) 1 110CT93 D 1 3 Sigambra tentaculata 1 110CT93 D 1 3 Paraprionospio pinnata 1 110CT93 D 1 3 Cossura delta 2 110CT93 D 1 3 Armandia maculata 1 110CT93 D 1 3 Mediomastus ambiseta 3 110CT93 D 1 10 Rhynchocoel (unidentified) 2 110CT93 D 1 10 Oligochaetes (unidentified) 1 110CT93 D 1 10 Lumbrineris parvapedata 1 110CT93 D 1 10 Minuspio cirrifera 2 110CT93 D 1 10 Cossura delta 3 110CT93 D 1 10 Ophiuroidea (unidentified) 1 110CT93 D 1 10 Apseudes sp. A 1 110CT93 D 1 10 Naineris sp. A 1 110CT93 D 1 10 Mediomastus ambiseta 2 110CT93 D 2 3 Rhynchocoel (unidentified) 2 110CT93 D 2 3 Streblospio benedicti 2 110CT93 D 2 3 Paraprionospio pinnata 1 110CT93 D 2 3 Minuspio cirrifera 3 110CT93 D 2 3 Periploma margaritaceum 4 110CT93 D 2 3 Ophiuroidea (unidentified) 1 110CT93 D 2 3 Mediomastus ambiseta 4 110CT93 D 2 10 Rhynchocoel (unidentified) 2 110CT93 D 2 10 Oligochaetes (unidentified) 1 110CT93 D 2 10 Paleanotus heteroseta 1 110CT93 D 2 10 Cossura delta 2 110CT93 D 2 10 Periploma margaritaceum 2 110CT93 D 2 10 Ophiuroidea (unidentified) 1 110CT93 D 2 10 Periploma cf. orbiculare 1 110CT93 D 2 10 Mediomastus ambiseta 2 110CT93 D 3 3 Minuspio cirrifera 2 110CT93 D 3 3 Periploma margaritaceum 5 110CT93 D 3 3 Mediomastus ambiseta 2 110CT93 D 3 10 Rhynchocoel (unidentified) 1 110CT93 D 3 10 Minuspio cirrifera 7 110CT93 D 3 10 Periploma margaritaceum 2 110CT93 D 3 10 Ophiuroidea (unidentified) 1 110CT93 D 3 10 Mediomastus ambiseta 1 110CT93 E 1 3 Streblospio benedicti 2 110CT93 E 1 3 Nassarius acutus 1 110CT93 E 1 3 Pyramidella crenulata 1 110CT93 E 1 3 Mediomastus ambiseta 8 110CT93 E 1 10 Oligochaetes (unidentified) 2 110CT93 E 1 10 Streblospio benedicti 2 110CT93 E 1 10 Paraprionospio pinnata 2 110CT93 E 1 10 Mediomastus ambiseta 3 110CT93 E 2 3 Streblospio benedicti 8 110CT93 E 2 3 Paraprionospio pinnata 1 110CT93 E 2 3 Pyramidella crenulata 1 110CT93 E 2 3 Mediomastus ambiseta 4 110CT93 E 2 10 Gyptis vittata 2 110CT93 E 2 10 Streblospio benedicti 1 110CT93 E 2 10 Paraprionospio pinnata 2 110CT93 E 2 10 Ancistrosyllis groenlandica 1 110CT93 E 2 10 Paraonidae Grp. A 1 110CT93 E 2 10 Caecum johnsoni 1 110CT93 E 2 10 Mediomastus ambiseta 5 110CT93 E 3 3 Streblospio benedicti 4 110CT93 E 3 3 Nassarius acutus 1 110CT93 E 3 3 Mediomastus ambiseta 3 110CT93 E 3 10 Paraprionospio pinnata 4 110CT93 E 3 10 Cossura delta 1 110CT93 E 3 10 Mediomastus ambiseta 2 110CT93 F 1 3 Streblospio benedicti 4 110CT93 F 1 3 Paraprionospio pinnata 1 110CT93 F 1 3 Ampelisca abdita 4 110CT93 F 1 3 Mediomastus ambiseta 1 110CT93 F 1 10 Streblospio benedicti 1 110CT93 F 1 10 Paraprionospio pinnata 1 110CT93 F 1 10 Parandalia ocularis 2 110CT93 F 1 10 Mediomastus ambiseta 1 110CT93 F 2 3 Rhynchocoel (unidentified) 1 110CT93 F 2 3 Streblospio benedicti 6 110CT93 F 2 3 Paraprionospio pinnata 1 110CT93 F 2 3 Ampelisca abdita 2 110CT93 F 2 3 Megalops 1 110CT93 F 2 3 Mediomastus ambiseta 5 110CT93 F 2 10 Cossura delta 1 110CT93 F 2 10 Mediomastus ambiseta 2 110CT93 F 3 3 Rhynchocoel (unidentified) 1 110CT93 F 3 3 Streblospio benedicti 3 110CT93 F 3 3 Spiochaetopterus costarum 1 11OCT03 F 3 3 Ampelisca abdita 1 110CT93 F 3 3 Nassarius acutus 1 110CT93 F 3 3 Mediomastus ambiseta 5 110CT93 F 3 10 No species observed 0 05JAN94 A 1 3 Polydora websteri 2 05JAN94 A 1 3 Streblospio benedicti 6 05JAN94 A 1 3 Haploscoloplos foliosus 1 05JAN94 A 1 3 Capitella capitata 3 05JAN94 A 1 3 Capitellides jonesi 2 05JAN94 A 1 3 Notomastus latericeus 1 05JAN94 A 1 3 Mulinia lateralis 2 05JAN94 A 1 3 Cyclaspis varians 1 05JAN94 A 1 3 Macoma mitchelli 1 05JAN94 A 1 3 Mediomastus ambiseta 14 05JAN94 A 1 10 Mediomastus ambiseta 1 05JAN94 A 2 3 Rhynchocoel (unidentified) 1 05JAN94 A 2 3 Streblospio benedicti 11 05JAN94 A 2 3 Capitella capitata 1 05JAN94 A 2 3 Mulinia lateralis 4 05JAN94 A 2 3 Cyclaspis varians 3 05JAN94 A 2 3 Macoma mitchelli 1 05JAN94 A 2 3 Mediomastus ambiseta 23 05JAN94 A 2 10 Mediomastus ambiseta 2 05JAN94 A 3 3 Streblospio benedicti 6 05JAN94 A 3 3 Mulinia lateralis 8 05JAN94 A 3 3 Brachidontes exustus 1 05JAN94 A 3 3 Macoma mitchelli 2 05JAN94 A 3 3 Mediomastus ambiseta 17 05JAN94 A 3 10 No species observed 0 05JAN94 B 1 3 Streblospio benedicti 5 05JAN94 B 1 3 Mulinia lateralis 5 05JAN94 B 1 3 Macoma mitchelli 2 05JAN94 B 1 3 Mediomastus ambiseta 2 05JAN94 B 1 10 Neanthes succinea 1 05JAN94 B 1 10 Ogyrides limicola 1 05JAN94 B 1 10 Mediomastus ambiseta 4 05JAN94 B 2 3 Streblospio benedicti 4 05JAN94 B 2 3 Haploscoloplos foliosus 2 05JAN94 B 2 3 Mulinia lateralis 5 05JAN94 B 2 3 Macoma mitchelli 1 05JAN94 B 2 3 Mediomastus ambiseta 5 05JAN94 B 2 10 Mediomastus ambiseta 1 05JAN94 B 3 3 Glycinde solitaria 1 05JAN94 B 3 3 Haploscoloplos foliosus 2 05JAN94 B 3 3 Mulinia lateralis 7 05JAN94 B 3 3 Mediomastus ambiseta 10 05JAN94 B 3 10 Paraprionospio pinnata 1 05JAN94 B 3 10 Capitellidae (unidentified) 1 05JAN94 B 3 10 Macoma mitchelli 2 05JAN94 B 3 10 Mediomastus ambiseta 2 05JAN94 C 1 3 Sigambra bassi 1 05JAN94 C 1 3 Paraprionospio pinnata 1 05JAN94 C 1 3 Haploscoloplos foliosus 1 05JAN94 C 1 3 Diastylis sp. 1 05JAN94 C 1 3 Mediomastus ambiseta 8 05JAN94 C 1 10 Rhynchocoel (unidentified) 1 05JAN94 C 1 10 Spiochaetopterus costarum 1 05JAN94 C 1 10 Mediomastus ambiseta 2 05JAN94 C 2 3 Rhynchocoel (unidentified) 2 05JAN94 C 2 3 Glycinde solitaria 1 05JAN94 C 2 3 Streblospio benedicti 1 05JAN94 C 2 3 Haploscoloplos foliosus 3 05JAN94 C 2 3 Nuculana acuta 1 05JAN94 C 2 3 Ophiuroidea (unidentified) 1 05JAN94 C 2 3 Mediomastus ambiseta 21 05JAN94 C 2 10 Rhynchocoel (unidentified) 1 05JAN94 C 2 10 Glycinde solitaria 1 05JAN94 C 2 10 Paraprionospio pinnata 1 05JAN94 C 2 10 Cossura delta 1 05JAN94 C 2 10 Mediomastus ambiseta 3 05JAN94 C 3 3 Rhynchocoel (unidentified) 2 05JAN94 C 3 3 Neanthes succinea 1 05JAN94 C 3 3 Glycinde solitaria 1 05JAN94 C 3 3 Haploscoloplos foliosus 3 05JAN94 C 3 3 Polychaete juv. (unidentified) 1 05JAN94 C 3 3 Mediomastus ambiseta 15 05JAN94 C 3 10 Streblospio benedicti 1 05JAN94 C 3 10 Maldanidae (unidentified) 1 05JAN94 C 3 10 Polychaete juv. (unidentified) 1 05JAN94 C 3 10 Mediomastus ambiseta 1 05JAN94 D 1 3 Rhynchocoel (unidentified) 1 05JAN94 D 1 3 Podarke obscura 1 05JAN94 D 1 3 Glycinde solitaria 1 05JAN94 D 1 3 Diopatra cuprea 1 05JAN94 D 1 3 Paraprionospio pinnata 1 05JAN94 D 1 3 Mulinia lateralis 2 05JAN94 D 1 3 Ophiuroidea (unidentified) 1 05JAN94 D 1 3 Armandia maculata 1 05JAN94 D 1 3 Apseudes sp. A 2 05JAN94 D 1 3 Mediomastus ambiseta 16 05JAN94 D 1 10 Ophiuroidea (unidentified) 1 05JAN94 D 2 3 Anthozoa (unidentified) 1 05JAN94 D 2 3 Diopatra cuprea 2 05JAN94 D 2 3 Lumbrineris latreilli 1 05JAN94 D 2 3 Minuspio cirrifera 2 05JAN94 D 2 3 Cossura delta 1 05JAN94 D 2 3 Phoronis architecta 1 05JAN94 D 2 3 Listriella barnardi 1 05JAN94 D 2 3 Ophiuroidea (unidentified) 1 05JAN94 D 2 3 Hemicyclops sp. 1 05JAN94 D 2 3 Turbellaria (unidentified) 1 05JAN94 D 2 3 Apseudes sp. A 2 05JAN94 D 2 3 Mediomastus ambiseta 15 05JAN94 D 2 10 Rhynchocoel (unidentified) 2 05JAN94 D 2 10 Minuspio cirrifera 7 05JAN94 D 2 10 Mulinia lateralis 1 05JAN94 D 2 10 Lumbrineris tenuis 1 05JAN94 D 2 10 Ophiuroidea (unidentified) 1 05JAN94 D 2 10 Mediomastus ambiseta 10 05JAN94 D 3 3 Anthozoa (unidentified) 2 05JAN94 D 3 3 Rhynchocoel (unidentified) 2 05JAN94 D 3 3 Eteone heteropoda 1 05JAN94 D 3 3 Glycinde solitaria 1 05JAN94 D 3 3 Lumbrineris latreilli 3 05JAN94 D 3 3 Paraprionospio pinnata 2 05JAN94 D 3 3 Minuspio cirrifera 5 05JAN94 D 3 3 Notomastus latericeus 1 05JAN94 D 3 3 Phoronis architecta 1 05JAN94 D 3 3 Ophiuroidea (unidentified) 1 05JAN94 D 3 3 Pelecypoda (unidentified) 1 05JAN94 D 3 3 Turbellaria (unidentified) 1 05JAN94 D 3 3 Mediomastus ambiseta 13 05JAN94 D 3 3 Eudorella sp. 1 05JAN94 D 3 10 Rhynchocoel (unidentified) 2 05JAN94 D 3 10 Podarke obscura 2 05JAN94 D 3 10 Minuspio cirrifera 2 05JAN94 D 3 10 Haploscoloplos foliosus 1 05JAN94 D 3 10 Cossura delta 3 05JAN94 D 3 10 Mulinia lateralis 5 05JAN94 D 3 10 Ophiuroidea (unidentified) 1 05JAN94 D 3 10 Macoma mitchelli 2 05JAN94 D 3 10 Apseudes sp. A 4 05JAN94 D 3 10 Mediomastus ambiseta 13 05JAN94 E 1 3 Podarke obscura 2 05JAN94 E 1 3 Glycinde solitaria 1 05JAN94 E 1 3 Streblospio benedicti 1 05JAN94 E 1 3 Paraprionospio pinnata 2 05JAN94 E 1 3 Minuspio cirrifera 1 05JAN94 E 1 3 Cossura delta 2 05JAN94 E 1 3 Nuculana acuta 1 05JAN94 E 1 3 Mulinia lateralis 68 05JAN94 E 1 3 Acteocina canaliculata 1 05JAN94 E 1 3 Ophiuroidea (unidentified) 1 05JAN94 E 1 3 Pyramidella crenulata 3 05JAN94 E 1 3 Mediomastus ambiseta 32 05JAN94 E 1 10 Gyptis vittata 1 05JAN94 E 1 10 Paraprionospio pinnata 2 05JAN94 E 1 10 Apoprionospio pygmaea 1 05JAN94 E 1 10 Haploscoloplos foliosus 4 05JAN94 E 1 10 Cossura delta 4 05JAN94 E 1 10 Paraonidae Grp. A 1 05JAN94 E 1 10 Mediomastus ambiseta 9 05JAN94 E 1 10 Glycinde nordmanni 1 05JAN94 E 2 3 Streblospio benedicti 4 05JAN94 E 2 3 Cossura delta 2 05JAN94 E 2 3 Haminoea succinea 1 05JAN94 E 2 3 Nuculana acuta 1 05JAN94 E 2 3 Mulinia lateralis 71 05JAN94 E 2 3 Acteocina canaliculata 2 05JAN94 E 2 3 Microprotopus spp. 1 05JAN94 E 2 3 Pyramidella crenulata 1 05JAN94 E 2 3 Turbellaria (unidentified) 1 05JAN94 E 2 3 Oxyurostylis smithi 1 05JAN94 E 2 3 Mediomastus ambiseta 11 05JAN94 E 2 10 Rhynchocoel (unidentified) 1 05JAN94 E 2 10 Podarke obscura 1 05JAN94 E 2 10 Paraprionospio pinnata 8 05JAN94 E 2 10 Minuspio cirrifera 1 05JAN94 E 2 10 Haploscoloplos foliosus 1 05JAN94 E 2 10 Mulinia lateralis 1 05JAN94 E 2 10 Hobsonia florida 1 05JAN94 E 2 10 Mediomastus ambiseta 14 05JAN94 E 3 3 Glycinde solitaria 1 05JAN94 E 3 3 Streblospio benedicti 1 05JAN94 E 3 3 Paraprionospio pinnata 2 05JAN94 E 3 3 Cossura delta 3 05JAN94 E 3 3 Nuculana acuta 4 05JAN94 E 3 3 Mulinia lateralis 96 05JAN94 E 3 3 Balanus eburneus 1 05JAN94 E 3 3 Acteocina canaliculata 5 05JAN94 E 3 3 Ophiuroidea (unidentified) 1 05JAN94 E 3 3 Pyramidella crenulata 1 05JAN94 E 3 3 Turbellaria (unidentified) 2 05JAN94 E 3 3 Mediomastus ambiseta 15 05JAN94 E 3 10 Podarke obscura 3 05JAN94 E 3 10 Paraprionospio pinnata 5 05JAN94 E 3 10 Haploscoloplos foliosus 1 05JAN94 E 3 10 Cossura delta 3 05JAN94 E 3 10 Haminoea succinea 1 05JAN94 E 3 10 Acteocina canaliculata 1 05JAN94 E 3 10 Mediomastus ambiseta 24 05JAN94 F 1 3 Streblospio benedicti 7 05JAN94 F 1 3 Mulinia lateralis 1 05JAN94 F 1 3 Ogyrides limicola 1 05JAN94 F 1 3 Phoronis architecta 1 05JAN94 F 1 3 Pelecypoda (unidentified) 1 05JAN94 F 1 3 Macoma mitchelli 8 05JAN94 F 1 3 Mediomastus ambiseta 5 05JAN94 F 1 10 Phoronis architecta 1 05JAN94 F 2 3 Rhynchocoel (unidentified) 1 05JAN94 F 2 3 Streblospio benedicti 5 05JAN94 F 2 3 Flaploscoloplos foliosus 1 05JAN94 F 2 3 Mulinia lateralis 1 05JAN94 F 2 3 Ampelisca abdita 1 05JAN94 F 2 3 Acteocina canaliculata 1 05JAN94 F 2 3 Macoma mitchelli 11 05JAN94 F 2 3 Mediomastus ambiseta 9 05JAN94 F 2 3 Glycinde nordmanni 1 05JAN94 F 2 10 Paraprionospio pinnata 1 05JAN94 F 3 3 Podarke obscura 1 05JAN94 F 3 3 Streblospio benedicti 4 05JAN94 F 3 3 Haploscoloplos foliosus 2 05JAN94 F 3 3 Mulinia lateralis 1 05JAN94 F 3 3 Acteocina canaliculata 2 05JAN94 F 3 3 Macoma mitchelli 13 05JAN94 F 3 3 Mediomastus ambiseta 7 05JAN94 F 3 10 Rhynchocoel (unidentified) 1 05JAN94 F 3 10 Haploscoloplos foliosus 1 05JAN94 F 3 10 Cossura delta 1 05JAN94 F 3 10 Acteocina canaliculata 2 05JAN94 F 3 10 Macoma mitchelli 5 05JAN94 F 3 10 Mediomastus ambiseta 1 07APR94 A 1 3 Eteone heteropoda 2 07APR94 A 1 3 Streblospio benedicti 7 07APR94 A 1 3 Notomastus latericeus 1 07APR94 A 1 3 Nuculana acuta 1 07APR94 A 1 3 Mulinia lateralis 7 07APR94 A 1 3 Ampelisca abdita 1 07APR94 A 1 3 Mediomastus ambiseta 13 07APR94 A 1 3 Glycinde nordmanni 1 07APR94 A 1 10 Rhynchocoel (unidentified) 1 07APR94 A 1 10 Mediomastus ambiseta 7 07APR94 A 2 3 Streblospio benedicti 11 07APR94 A 2 3 Mulinia lateralis 9 07APR94 A 2 3 Cyclaspis varians 2 07APR94 A 2 3 Mediomastus ambiseta 14 07APR94 A 2 10 Mysidopsis sp. 1 07APR94 A 2 10 Macoma mitchelli 1 07APR94 A 2 10 Mediomastus ambiseta 14 07APR94 A 3 3 Oligochaetes (unidentified) 4 07APR94 A 3 3 Streblospio benedicti 15 07APR94 A 3 3 Capitella capitata 1 07APR94 A 3 3 Mulinia lateralis 10 07APR94 A 3 3 Cyclaspis varians 1 07APR94 A 3 3 Mediomastus ambiseta 18 07APR94 A 3 10 Macoma mitchelli 1 07APR94 A 3 10 Mediomastus ambiseta 18 07APR94 A 3 10 Glycinde nordmanni 1 07APR94 B 1 3 Cossura delta 1 07APR94 B 1 3 Turbellaria (unidentified) 1 07APR94 B 1 3 Mediomastus ambiseta 26 07APR94 B 1 3 Glycinde nordmanni 4 07APR94 B 1 10 Haploscoloplos foliosus 1 07APR94 B 1 10 Macoma mitchelli 1 07APR94 B 1 10 Mediomastus ambiseta 5 07APR94 B 2 3 Streblospio benedicti 4 07APR94 B 2 3 Mulinia lateralis 3 07APR94 B 2 3 Mediomastus ambiseta 20 07APR94 B 2 3 Glycinde nordmanni 4 07APR94 B 2 10 Rhynchocoel (unidentified) 1 07APR94 B 2 10 Flaploscoloplos foliosus 1 07APR94 B 2 10 Parandalia ocularis 1 07APR94 B 2 10 Mediomastus ambiseta 1 07APR94 B 3 3 Rhynchocoel (unidentified) 1 07APR94 B 3 3 Streblospio benedicti 3 07APR94 B 3 3 Mulinia lateralis 4 07APR94 B 3 3 Nereidae (unidentified) 1 07APR94 B 3 3 Mediomastus ambiseta 9 07APR94 B 3 10 Rhynchocoel (unidentified) 1 07APR94 B 3 10 Haploscoloplos foliosus 1 07APR94 B 3 10 Mediomastus ambiseta 3 07APR94 C 1 3 Rhynchocoel (unidentified) 1 07APR94 C 1 3 Podarke obscura 1 07APR94 C 1 3 Haploscoloplos foliosus 2 07APR94 C 1 3 Mediomastus ambiseta 2 07APR94 C 1 3 Glycinde nordmanni 1 07APR94 C 1 10 Mediomastus ambiseta 2 07APR94 C 1 10 Glycinde nordmanni 1 07APR94 C 2 3 Gyptis vittata 1 07APR94 C 2 3 Haploscoloplos foliosus 2 07APR94 C 2 3 Mediomastus ambiseta 1 07APR94 C 2 10 Paraonidae Grp. B 1 07APR94 C 2 10 Mediomastus ambiseta 6 07APR94 C 3 3 Paraonidae Grp. A 1 07APR94 C 3 3 Mediomastus ambiseta 3 07APR94 C 3 10 Sigambra tentaculata 1 07APR94 C 3 10 Haploscoloplos foliosus 1 07APR94 C 3 10 Mediomastus ambiseta 3 07APR94 C 3 10 Glycinde nordmanni 1 07APR94 D 1 3 Anthozoa (unidentified) 2 07APR94 D 1 3 Rhynchocoel (unidentified) 1 07APR94 D 1 3 Oligochaetes (unidentified) 1 07APR94 D 1 3 Glycera americana 1 07APR94 D 1 3 Minuspio cirrifera 3 07APR94 D 1 3 Haploscoloplos foliosus 1 07APR94 D 1 3 Cyclaspis varians 2 07APR94 D 1 3 Ampelisca abdita 1 07APR94 D 1 3 Sphaerosyllis sp. A 1 07APR94 D 1 3 Mediomastus ambiseta 11 07APR94 D 1 10 Rhynchocoel (unidentified) 2 07APR94 D 1 10 Oligochaetes (unidentified) 1 07APR94 D 1 10 Paleanotus heteroseta 1 07APR94 D 1 10 Minuspio cirrifera 21 07APR94 D 1 10 Cossura delta 1 07APR94 D 1 10 Ophiuroidea (unidentified) 1 07APR94 D 1 10 Naineris sp. A 3 07APR94 D 1 10 Mediomastus ambiseta 2 07APR94 D 2 3 Cossura delta 1 07APR94 D 2 3 Turbellaria (unidentified) 1 07APR94 D 2 3 Mediomastus ambiseta 9 07APR94 D 2 10 Oligochaetes (unidentified) 3 07APR94 D 2 10 Onuphis sp. 1 07APR94 D 2 10 Minuspio cirrifera 3 07APR94 D 2 10 Ophiuroidea (unidentified) 1 07APR94 D 2 10 Naineris sp. A 3 07APR94 D 2 10 Mediomastus ambiseta 3 07APR94 D 3 3 Rhynchocoel (unidentified) 1 07APR94 D 3 3 Oligochaetes (unidentified) 2 07APR94 D 3 3 Ancistrosyllis papillosa 1 07APR94 D 3 3 Minuspio cirrifera 2 07APR94 D 3 3 Cossura delta 3 07APR94 D 3 3 Ophiuroidea (unidentified) 1 07APR94 D 3 3 Apseudes sp. A 1 07APR94 D 3 3 Periploma cf. orbiculare 2 07APR94 D 3 3 Sigambra cf. wassi 1 07APR94 D 3 3 Mediomastus ambiseta 8 07APR94 D 3 10 Oligochaetes (unidentified) 1 07APR94 D 3 10 Diopatra cuprea 1 07APR94 D 3 10 Lumbrinehs parvapedata 1 07APR94 D 3 10 Minuspio cirrifera 7 07APR94 D 3 10 Cyclaspis varians 1 07APR94 D 3 10 Ophiuroidea (unidentified) 1 07APR94 D 3 10 Periploma cf. orbiculare 2 07APR94 D 3 10 Mediomastus ambiseta 2 07APR94 E 1 3 Cossura delta 2 07APR94 E 1 3 Nuculana acuta 7 07APR94 E 1 3 Mulinia lateralis 16 07APR94 E 1 3 Acteocina canaliculata 6 07APR94 E 1 3 Ophiuroidea (unidentified) 1 07APR94 E 1 3 Eulimostoma sp. 1 07APR94 E 1 3 Mediomastus ambiseta 16 07APR94 E 1 10 Podarke obscura 1 07APR94 E 1 10 Paraprionospio pinnata 3 07APR94 E 1 10 Cossura delta 4 07APR94 E 1 10 Nuculana acuta 1 07APR94 E 1 10 Ophiuroidea (unidentified) 2 07APR94 E 1 10 Mediomastus ambiseta 13 07APR94 E 2 3 Eteone heteropoda 2 07APR94 E 2 3 Streblospio benedicti 1 07APR94 E 2 3 Paraprionospio pinnata 2 07APR94 E 2 3 Cossura delta 4 07APR94 E 2 3 Balanus eburneus 3 07APR94 E 2 3 Eulimostoma sp. 1 07APR94 E 2 3 Asychis elongata 1 07APR94 E 2 3 Oxyurostylis smithi 1 07APR94 E 2 3 Mediomastus ambiseta 16 07APR94 E 2 10 Rhynchocoel (unidentified) 1 07APR94 E 2 10 Oligochaetes (unidentified) 1 07APR94 E 2 10 Podarke obscura 2 07APR94 E 2 10 Paraprionospio pinnata 3 07APR94 E 2 10 Scolelepis texana 1 07APR94 E 2 10 Haploscoloplos foliosus 1 07APR94 E 2 10 Cossura delta 5 07APR94 E 2 10 Nuculana acuta 1 07APR94 E 2 10 Mediomastus ambiseta 11 07APR94 E 2 10 Glycinde nordmanni 1 07APR94 E 3 3 Anthozoa (unidentified) 1 07APR94 E 3 3 Oligochaetes (unidentified) 1 07APR94 E 3 3 Podarke obscura 2 07APR94 E 3 3 Streblospio benedicti 1 07APR94 E 3 3 Cossura delta 7 07APR94 E 3 3 Nuculana acuta 5 07APR94 E 3 3 Mulinia lateralis 22 07APR94 E 3 3 Balanus eburneus 6 07APR94 E 3 3 Acteocina canaliculata 7 07APR94 E 3 3 Ophiuroidea (unidentified) 1 07APR94 E 3 3 Asychis elongata 1 07APR94 E 3 3 Mediomastus ambiseta 22 07APR94 E 3 10 Lumbrineris parvapedata 2 07APR94 E 3 10 Paraprionospio pinnata 3 07APR94 E 3 10 Haploscoloplos foliosus 2 07APR94 E 3 10 Cossura delta 3 07APR94 E 3 10 Paraonidae Grp. A 2 07APR94 E 3 10 Mediomastus ambiseta 12 07APR94 E 3 10 Glycinde nordmanni 1 07APR94 F 1 3 Streblospio benedicti 3 07APR94 F 1 3 Haploscoloplos foliosus 1 07APR94 F 1 3 Mulinia lateralis 1 07APR94 F 1 3 Corophium louisianum 1 07APR94 F 1 3 Acteocina canaliculata 2 07APR94 F 1 3 Macoma mitchelli 1 07APR94 F 1 3 Mediomastus ambiseta 4 07APR94 F 1 10 Macoma mitchelli 1 07APR94 F 2 3 Streblospio benedicti 2 07APR94 F 2 3 Mulinia lateralis 2 07APR94 F 2 3 Cyclaspis varians 2 07APR94 F 2 3 Acteocina canaliculata 1 07APR94 F 2 3 Macoma mitchelli 1 07APR94 F 2 3 Mediomastus ambiseta 7 07APR94 F 2 10 Gyptis vittata 2 07APR94 F 2 10 Paraprionospio pinnata 1 07APR94 F 2 10 Flaploscoloplos foliosus 1 07APR94 F 2 10 Caecum pulchellum 1 07APR94 F 2 10 Macoma mitchelli 2 07APR94 F 2 10 Parandalia ocularis 1 07APR94 F 3 3 Streblospio benedicti 1 07APR94 F 3 3 Paraprionospio pinnata 1 07APR94 F 3 3 Mulinia lateralis 3 07APR94 F 3 3 Corophium louisianum 5 07APR94 F 3 3 Mediomastus ambiseta 3 07APR94 F 3 10 Paraprionospio pinnata 1 07APR94 F 3 10 Haploscoloplos foliosus 1 07APR94 F 3 10 Cossura delta 1 07APR94 F 3 10 Glycinde nordmanni 1 Guadalupe Estuary 070CT92 A 1 3 Streblospio benedicti 1 070CT92 A 1 3 Mulinia lateralis 8 070CT92 A 1 3 Chironomid larvae 1 070CT92 A 1 3 Hobsonia florida 1 070CT92 A 1 3 Littoridina sphinctostoma 74 070CT92 A 1 10 Littoridina sphinctostoma 1 070CT92 A 2 3 Mulinia lateralis 3 070CT92 A 2 3 Chironomid larvae 2 070CT92 A 2 3 Hobsonia florida 2 070CT92 A 2 3 Littoridina sphinctostoma 12 070CT92 A 2 10 Oligochaetes (unidentified) 1 070CT92 A 2 10 Chironomid larvae 1 070CT92 A 2 10 Hobsonia florida 2 070CT92 A 3 3 Streblospio benedicti 1 070CT92 A 3 3 Mulinia lateralis 9 070CT92 A 3 3 Hobsonia florida 3 070CT92 A 3 3 Rangia cuneata 1 070CT92 A 3 3 Littoridina sphinctostoma 106 070CT92 A 3 10 No species observed 0 070CT92 B 1 3 Streblospio benedicti 74 070CT92 B 1 3 Mulinia lateralis 1 070CT92 B 1 3 Mediomastus ambiseta 9 070CT92 B 1 10 Streblospio benedicti 4 070CT92 B 1 10 Mediomastus ambiseta 19 070CT92 B 2 3 Streblospio benedicti 65 070CT92 B 2 3 Mediomastus ambiseta 9 070CT92 B 2 10 Rhynchocoel (unidentified) 1 070CT92 B 2 10 Streblospio benedicti 4 070CT92 B 2 10 Chironomid larvae 1 070CT92 B 2 10 Hobsonia florida 1 070CT92 B 2 10 Mediomastus ambiseta 19 070CT92 B 3 3 Streblospio benedicti 86 070CT92 B 3 3 Mulinia lateralis 3 070CT92 B 3 3 Paraonidae Grp. B 1 070CT92 B 3 3 Hobsonia florida 1 070CT92 B 3 3 Mediomastus ambiseta 22 070CT92 B 3 10 Rhynchocoel (unidentified) 1 070CT92 B 3 10 Streblospio benedicti 1 070CT92 B 3 10 Capitella capitata 2 070CT92 B 3 10 Mediomastus ambiseta 19 070CT92 C 1 3 Streblospio benedicti 11 070CT92 C 1 3 Mediomastus ambiseta 16 070CT92 C 1 10 Rhynchocoel (unidentified) 2 070CT92 C 1 10 Callianassa sp. 1 070CT92 C 1 10 Parandalia ocularis 1 070CT92 C 1 10 Mediomastus ambiseta 3 070CT92 C 2 3 Streblospio benedicti 4 070CT92 C 2 3 Mediomastus ambiseta 14 070CT92 C 2 10 Rhynchocoel (unidentified) 5 070CT92 C 2 10 Callianassa sp. 1 070CT92 C 2 10 Parandalia ocularis 2 070CT92 C 2 10 Mediomastus ambiseta 14 070CT92 C 3 3 Rhynchocoel (unidentified) 2 070CT92 C 3 3 Streblospio benedicti 13 070CT92 C 3 3 Mulinia lateralis 1 070CT92 C 3 3 Mediomastus ambiseta 16 070CT92 C 3 10 Rhynchocoel (unidentified) 2 070CT92 C 3 10 Streblospio benedicti 1 070CT92 C 3 10 Mediomastus ambiseta 6 070CT92 D 1 3 Glycinde solitaria 1 070CT92 D 1 3 Streblospio benedicti 1 070CT92, D 1 3 Mulinia lateralis 1 070CT92 D 1 3 Ogyrides limicola 1 070CT92 D 1 3 Mediomastus ambiseta 5 070CT92 D 1 10 Parandalia ocularis 2 070CT92 D 1 10 Mediomastus ambiseta 1 070CT92 D 2 3 Rhynchocoel (unidentified) 2 070CT92 D 2 3 Streblospio benedicti 1 070CT92 D 2 3 Paraprionospio pinnata 1 070CT92 D 2 3 Monoculodes sp. 1 070CT92 D 2 3 Littoridina sphinctostoma 1 070CT92 D 2 3 Mediomastus ambiseta 10 070CT92 D 2 10 Thompsonula sp. 1 070CT92 D 2 10 Mediomastus ambiseta 3 070CT92 D 3 3 Rhynchocoel (unidentified) 1 070CT92 D 3 3 Streblospio benedicti 1 070CT92 D 3 3 Littoridina sphinctostoma 1 070CT92 D 3 3 Mediomastus ambiseta 7 070CT92 D 3 10 Mediomastus ambiseta 3 12JAN93 A 1 3 Rhynchocoel (unidentified) 2 12JAN93 A 1 3 Streblospio benedicti 2 12JAN93 A 1 3 Mulinia lateralis 13 12JAN93 A 1 3 Littoridina sphinctostoma 125 12JAN93 A 1 3 Mediomastus ambiseta 4 12JAN93 A 1 10 Chironomid larvae 1 12JAN93 A 1 10 Mediomastus ambiseta 1 12JAN93 A 2 3 Rhynchocoel (unidentified) 1 12JAN93 A 2 3 Streblospio benedicti 3 12JAN93 A 2 3 Mulinia lateralis 9 12JAN93 A 2 3 Littoridina sphinctostoma 147 12JAN93 A 2 3 Mediomastus ambiseta 7 12JAN93 A 2 10 Rhynchocoel (unidentified) 1 12JAN93 A 2 10 Pieces 1 12JAN93 A 3 3 Rhynchocoel (unidentified) 1 12JAN93 A 3 3 Eteone heteropoda 1 12JAN93 A 3 3 Streblospio benedicti 3 12JAN93 A 3 3 Mulinia lateralis 17 12JAN93 A 3 3 Monoculodes sp. 1 12JAN93 A 3 3 Littoridina sphinctostoma 117 12JAN93 A 3 3 Mediomastus ambiseta 7 12JAN93 A 3 10 Capitella capitata 1 12JAN93 B 1 3 Rhynchocoel (unidentified) 1 12JAN93 B 1 3 Diopatra cuprea 1 12JAN93 B 1 3 Streblospio benedicti 4 12JAN93 B 1 3 Capitella capitata 1 12JAN93 B 1 3 Mulinia lateralis 53 12JAN93 B 1 3 Brachidontes exustus 1 12JAN93 B 1 3 Macoma mitchelli 1 12JAN93 B 1 3 Mediomastus ambiseta 27 12JAN93 B 1 10 Capitella capitata 2 12JAN93 B 1 10 Mediomastus ambiseta 3 12JAN93 B 2 3 Streblospio benedicti 7 12JAN93 B 2 3 Capitella capitata 2 12JAN93 B 2 3 Mulinia lateralis 85 12JAN93 B 2 3 Macoma mitchelli 2 12JAN93 B 2 3 Mediomastus ambiseta 32 12JAN93 B 2 10 Mulinia lateralis 1 12JAN93 B 2 10 Mediomastus ambiseta 1 12JAN93 B 3 3 Rhynchocoel (unidentified) 1 12JAN93 B 3 3 Streblospio benedicti 3 12JAN93 B 3 3 Mulinia lateralis 63 12JAN93 B 3 3 Mediomastus ambiseta 15 12JAN93 B 3 10 Rhynchocoel (unidentified) 1 12JAN93 B 3 10 Glycinde solitaria 1 12JAN93 B 3 10 Capitella capitata 2 12JAN93 B 3 10 Mediomastus ambiseta 7 12JAN93 C 1 3 Glycinde solitaria 1 12JAN93 C 1 3 Lysidice ninetta 35 12JAN93 C 1 3 Streblospio benedicti 7 12JAN93 C 1 3 Mulinia lateralis 5 12JAN93 C 1 3 Monoculodes sp. 2 12JAN93 C 1 3 Parandalia ocularis 1 12JAN93 C 1 10 Rhynchocoel (unidentified) 2 12JAN93 C 1 10 Lysidice ninetta 5 12JAN93 C 1 10 Parandalia ocularis 1 12JAN93 C 2 3 Glycinde solitaria 1 12JAN93 C 2 3 Lysidice ninetta 22 12JAN93 C 2 3 Streblospio benedicti 2 12JAN93 C 2 3 Mulinia lateralis 1 12JAN93 C 2 3 Monoculodes sp. 4 12JAN93 C 2 10 Lysidice ninetta 3 12JAN93 C 2 10 Parandalia ocularis 2 12JAN93 C 3 3 Rhynchocoel (unidentified) 1 12JAN93 C 3 3 Lysidice ninetta 27 12JAN93 C 3 3 Streblospio benedicti 2 12JAN93 C 3 3 Mulinia lateralis 2 12JAN93 C 3 3 Monoculodes sp. 3 12JAN93 C 3 3 Pyramidella crenulata 1 12JAN93 C 3 3 Macoma mitchelli 2 12JAN93 C 3 10 Lysidice ninetta 1 12JAN93 D 1 3 Streblospio benedicti 7 12JAN93 D 1 3 Paraprionospio pinnata 1 12JAN93 D 1 3 Mulinia lateralis 14 12JAN93 D 1 3 Macoma mitchelli 1 12JAN93 D 1 3 Mediomastus ambiseta 29 12JAN93 D 1 10 Mediomastus ambiseta 1 12JAN93 D 2 3 Glycinde solitaria 5 12JAN93 D 2 3 Streblospio benedicti 3 12JAN93 D 2 3 Mulinia lateralis 41 12JAN93 D 2 3 Acteocina canaliculata 1 12JAN93 D 2 3 Molgula manhattensis 1 12JAN93 D 2 3 Mediomastus ambiseta 23 12JAN93 D 2 10 Rhynchocoel (unidentified) 1 12JAN93 D 2 10 Mulinia lateralis 2 12JAN93 D 2 10 Mediomastus ambiseta 2 12JAN93 D 3 3 Rhynchocoel (unidentified) 3 12JAN93 D 3 3 Diopatra cuprea 1 12JAN93 D 3 3 Mulinia lateralis 67 12JAN93 D 3 3 Macoma mitchelli 1 12JAN93 D 3 3 Hobsonia florida 1 12JAN93 D 3 3 Mediomastus ambiseta 15 12JAN93 D 3 10 Minuspio cirrifera 1 12JAN93 D 3 10 Mediomastus ambiseta 2 05APR93 A 1 3 Capitella capitata 1 05APR93 A 1 3 Mulinia lateralis 112 05APR93 A 1 3 Chironomid larvae 1 05APR93 A 1 3 Hobsonia florida 1 05APR93 A 1 3 Littoridina sphinctostoma 50 05APR93 A 13 Mediomastus ambiseta 9 05APR93 A 1 10 Oligochaetes (unidentified) 1 05APR93 A 1 10 Mulinia lateralis 2 05APR93 A 1 10 Littoridina sphinctostoma 1 05APR93 A 1 10 Mediomastus ambiseta 1 05APR93 A 2 3 Rhynchocoel (unidentified) 1 05APR93 A 2 3 Streblospio benedicti 7 05APR93 A 2 3 Capitella capitata 1 05APR93 A 2 3 Mulinia lateralis 101 05APR93 A 2 3 Hobsonia florida 1 05APR93 A 2 3 Littoridina sphinctostoma 79 05APR93 A 2 3 Mediomastus ambiseta 11 05APR93 A 2 10 Oligochaetes (unidentified) 1 05APR93 A 2 10 Littoridina sphinctostoma 1 05APR93 A 2 10 Mediomastus ambiseta 2 05APR93 A 3 3 Mulinia lateralis 70 05APR93 A 3 3 Hobsonia florida 1 05APR93 A 3 3 Littoridina sphinctostoma 50 05APR93 A 3 3 Mediomastus ambiseta 7 05APR93 A 3 10 Rhynchocoel (unidentified) 1 05APR93 A 3 10 Capitella capitate 1 05APR93 A 3 10 Mulinia lateralis 4 05APR93 A 3 10 Littoridina sphinctostoma 1 05APR93 A 3 10 Mediomastus ambiseta 2 05APR93 B 1 3 Streblospio benedicti 47 05APR93 B 1 3 Capitella capitata 4 05APR93 B 1 3 Mulinia lateralis 106 05APR93 B 1 3 Littoridina sphinctostoma 4 05APR93 B 1 3 Mediomastus ambiseta 68 05APR93 B 1 10 Capitella capitata 1 05APR93 B 1 10 Macoma mitchelli 1 05APR93 B 1 10 Mediomastus ambiseta 13 05APR93 B 2 3 Streblospio benedicti 52 05APR93 B 2 3 Capitella capitata 1 05APR93 B 2 3 Mulinia lateralis 41 05APR93 B 2 3 Mediomastus ambiseta 88 05APR93 B 2 10 Streblospio benedicti 2 05APR93 B 2 10 Capitella capitata 1 05APR93 B 2 10 Mulinia lateralis 3 05APR93 B 2 10 Mediomastus ambiseta 13 05APR93 B 3 3 Eteone heteropoda 4 05APR93 B 3 3 Neanthes succinea 1 05APR93 B 3 3 Streblospio benedicti 36 05APR93 B 3 3 Capitella capitata 1 05APR93 B 3 3 Mulinia lateralis 100 05APR93 B 3 3 Mediomastus ambiseta 49 05APR93 B 3 10 Rhynchocoel (unidentified) 1 05APR93 B 3 10 Gyptis vittata 2 05APR93 B 3 10 Capitella capitata 1 05APR93 B 3 10 Mulinia lateralis 2 05APR93 B 3 10 Littoridina sphinctostoma 1 05APR93 B 3 10 Mediomastus ambiseta 11 05APR93 C 1 3 Glycinde solitaria 1 05APR93 C 1 3 Streblospio benedicti 15 05APR93 C 1 3 Capitella capitata 4 05APR93 C 1 3 Mulinia lateralis 67 05APR93 C 1 3 Littoridina sphinctostoma 28 05APR93 C 1 3 Mediomastus ambiseta 48 05APR93 C 1 10 Capitella capitata 1 05APR93 C 1 10 Mediomastus ambiseta 3 05APR93 C 2 3 Eteone heteropoda 1 05APR93 C 2 3 Streblospio benedicti 32 05APR93 C 2 3 Capitella capitata 4 05APR93 C 2 3 Pectinaria gouldii 2 05APR93 C 2 3 Mulinia lateralis 100 05APR93 C 2 3 Littoridina sphinctostoma 60 05APR93 C 2 3 Mediomastus ambiseta 59 05APR93 C 2 10 Mediomastus ambiseta 1 05APR93 C 3 3 Rhynchocoel (unidentified) 1 05APR93 C 3 3 Streblospio benedicti 10 05APR93 C 3 3 Capitella capitata 2 05APR93 C 3 3 Pectinaria gouldii 2 05APR93 C 3 3 Mulinia lateralis 35 05APR93 C 3 3 Littoridina sphinctostoma 8 05APR93 C 3 3 Mediomastus ambiseta 49 05APR93 C 3 10 Rhynchocoel (unidentified) 2 05APR93 C 3 10 Parandalia ocularis 1 05APR93 C 3 10 Mediomastus ambiseta 14 05APR93 D 1 3 Glycinde solitaria 3 05APR93 D 1 3 Streblospio benedicti 3 05APR93 D 1 3 Mulinia lateralis 32 05APR93 D 1 3 Littoridina sphinctostoma 7 05APR93 D 1 3 Parandalia ocularis 1 05APR93 D 1 3 Mediomastus ambiseta 29 05APR93 D 1 10 Rhynchocoel (unidentified) 1 05APR93 D 1 10 Glycinde solitaria 1 05APR93 D 1 10 Mulinia lateralis 1 05APR93 D 1 10 Mediomastus ambiseta 10 05APR93 D 2 3 Rhynchocoel (unidentified) 1 05APR93 D 2 3 Streblospio benedicti 4 05APR93 D 2 3 Mulinia lateralis 23 05APR93 D 2 3 Macoma mitchelli 1 05APR93 D 2 3 Littoridina sphinctostoma 7 05APR93 D 2 3 Oxyurostylis sp. 1 05APR93 D 2 3 Mediomastus ambiseta 32 05APR93 D 2 10 Spiochaetopterus costarum 1 05APR93 D 2 10 Littoridina sphinctostoma 3 05APR93 D 2 10 Thompsonula sp. 1 05APR93 D 3 3 Streblospio benedicti 8 05APR93 D 3 3 Mulinia lateralis 17 05APR93 D 3 3 Littoridina sphinctostoma 8 05APR93 D 3 3 Oxyurostylis sp. 3 05APR93 D 3 3 Mediomastus ambiseta 50 05APR93 D 3 10 Glycinde solitaria 1 05APR93 D 3 10 Polydora caulleryi 1 05APR93 D 3 10 Mediomastus ambiseta 4 09JUL93 A 1 3 Mulinia lateralis 39 09JUL93 A 1 3 Chironomid larvae 3 09JUL93 A 1 3 Rangia cuneata 7 09JUL93 A 1 3 Littoridina sphinctostoma 3 09JUL93 A 1 3 Mediomastus ambiseta 7 09JUL93 A 1 10 Mediomastus ambiseta 3 09JUL93 A 2 3 Oligochaetes (unidentified) 1 09JUL93 A 2 3 Mulinia lateralis 48 09JUL93 A 2 3 Rangia cuneata 20 09JUL93 A 2 3 Littoridina sphinctostoma 32 09JUL93 A 2 3 Mediomastus ambiseta 3 09JUL93 A 2 10 Mulinia lateralis 1 09JUL93 A 3 3 Mulinia lateralis 74 09JUL93 A 3 3 Rangia cuneata 10 09JUL93 A 3 3 Littoridina sphinctostoma 7 09JUL93 A 3 3 Mediomastus ambiseta 8 09JUL93 A 3 10 Oligochaetes (unidentified) 1 09JUL93 B 1 3 Mulinia lateralis 70 09JUL93 B 1 3 Mediomastus ambiseta 2 09JUL93 B 1 10 Capitella capitata 3 09JUL93 B 1 10 Mulinia lateralis 3 09JUL93 B 1 10 Littoridina sphinctostoma 2 09JUL93 B 1 10 Mediomastus ambiseta 6 09JUL93 B 2 3 Mulinia lateralis 38 09JUL93 B 2 3 Littoridina sphinctostoma 1 09JUL93 B 2 3 Mediomastus ambiseta 5 09JUL93 B 2 10 Neanthes succinea 1 09JUL93 B 2 10 Capitella capitata 2 09JUL93 B 2 10 Mulinia lateralis 5 09JUL93 B 2 10 Littoridina sphinctostoma 2 09JUL93 B 2 10 Mediomastus ambiseta 19 09JUL93 B 3 3 Neanthes succinea 1 09JUL93 B 3 3 Mulinia lateralis 54 09JUL93 B 3 3 Littoridina sphinctostoma 1 09JUL93 B 3 3 Mediomastus ambiseta 7 09JUL93 B 3 10 Mulinia lateralis 1 09JUL93 B 3 10 Mediomastus ambiseta 21 09JUL93 C 1 3 Littoridina sphinctostoma 2 09JUL93 C 1 3 Mediomastus ambiseta 4 09JUL93 C 1 10 Mediomastus ambiseta 17 09JUL93 C 2 3 Streblospio benedicti 5 09JUL93 C 2 3 Mulinia lateralis 9 09JUL93 C 2 3 Monoculodes sp. 1 09JUL93 C 2 3 Littoridina sphinctostoma 4 09JUL93 C 2 3 Mediomastus ambiseta 6 09JUL93 C 2 10 Streblospio benedicti 1 09JUL93 C 2 10 Capitella capitata 1 09JUL93 C 2 10 Mulinia lateralis 2 09JUL93 C 2 10 Littoridina sphinctostoma 1 09JUL93 C 2 10 Mediomastus ambiseta 31 09JUL93 C 3 3 Streblospio benedicti 3 09JUL93 C 3 3 Capitella capitata 1 09JUL93 C 3 3 Mulinia lateralis 8 09JUL93 C 3 3 Littoridina sphinctostoma 18 09JUL93 C 3 3 Mediomastus ambiseta 12 09JUL93 C 3 10 Rhynchocoel (unidentified) 1 09JUL93 C 3 10 Capitella capitata 1 09JUL93 C 3 10 Mediomastus ambiseta 15 09JUL93 D 1 3 Mulinia lateralis 2 09JUL93 D 1 3 Littoridina sphinctostoma 1 09JUL93 D 1 3 Mediomastus ambiseta 5 09JUL93 D 1 10 Parandalia ocularis 3 09JUL93 D 1 10 Mediomastus ambiseta 7 09JUL93 D 2 3 Mediomastus ambiseta 8 09JUL93 D 2 10 No species observed 0 09JUL93 D 3 3 Streblospio benedicti 1 09JUL93 D 3 3 Mediomastus ambiseta 16 09JUL93 D 3 10 Mediomastus ambiseta 3 110CT93 A 1 3 Polydora websteri 4 110CT93 A 1 3 Streblospio benedicti 3 110CT93 A 1 3 Mulinia lateralis 16 110CT93 A 1 3 Hobsonia florida 2 110CT93 A 1 3 Rangia cuneata 5 110CT93 A 1 3 Littoridina sphinctostoma 4 110CT93 A 1 3 Mediomastus ambiseta 6 110CT93 A 1 10 No species observed 0 110CT93 A 2 3 Anthozoa (unidentified) 1 11CCT93 A 2 3 Polydora websteri 7 11CCT93 A 2 3 Streblospio benedicti 1 11CCT93 A 2 3 Mulinia lateralis 14 110CT93 A 2 3 Hobsonia florida 1 110CT93 A 2 3 Rangia cuneata 2 110CT93 A 2 3 Littoridina sphinctostoma 5 110CT93 A 2 3 Mediomastus ambiseta 1 110CT93 A 2 10 Mediomastus ambiseta 2 110CT93 A 3 3 Anthozoa (unidentified) 1 110CT93 A 3 3 Oligochaetes (unidentified) 1 110CT93 A 3 3 Mulinia lateralis 8 110CT93 A 3 3 Hobsonia florida 4 110CT93 A 3 3 Rangia cuneata 1 110CT93 A 3 3 Littoridina sphinctostoma 11 110CT93 A 3 3 Mediomastus ambiseta 4 110CT93 A 3 10 Mediomastus ambiseta 1 110CT93 B 1 3 Streblospio benedicti 1 110CT93 B 1 3 Muiinia lateralis 2 110CT93 B 1 3 Mediomastus ambiseta 2 110CT93 B 1 10 Mediomastus ambiseta 8 110CT93 B 2 3 Streblospio benedicti 5 110CT93 B 2 3 Muiinia lateralis 7 110CT93 B 2 3 Mediomastus ambiseta 2 110CT93 B 2 10 Mediomastus ambiseta 6 110CT93 B 3 3 Anthozoa (unidentified) 1 110CT93 B 3 3 Rhynchocoel (unidentified) 1 110CT93 B 3 3 Polydora websteri 1 110CT93 B 3 3 Streblospio benedicti 6 110CT93 B 3 3 Capitella capitata 2 110CT93 B 3 3 Muiinia lateralis 16 110CT93 B 3 3 Littoridina sphinctostoma 1 110CT93 B 3 3 Mediomastus ambiseta 2 110CT93 B 3 10 Mediomastus ambiseta 16 110CT93 C 1 3 Rhynchocoel (unidentified) 1 110CT93 C 1 3 Polydora websteri 1 110CT93 C 1 3 Streblospio benedicti 5 110CT93 C 1 3 Capitella capitata 1 110CT93 C 1 3 Muiinia lateralis 56 110CT93 C 1 3 Balanus eburneus 15 110CT93 C 1 3 Littoridina sphinctostoma 7 110CT93 C 1 3 Mediomastus ambiseta 8 110CT93 C 1 10 Streblospio benedicti 1 110CT93 C 1 10 Mediomastus ambiseta 5 110CT93 C 2 3 Streblospio benedicti 4 110CT93 C 2 3 Muiinia lateralis 3 110CT93 C 2 3 Turbellaria (unidentified) 1 11QCT93 C 2 3 Littoridina sphinctostoma 11 110CT93 C 2 3 Mediomastus ambiseta 7 110CT93 C 2 10 Mediomastus ambiseta 22 110CT93 C 3 3 Streblospio benedicti 3 110CT93 C 3 3 Muiinia lateralis 3 110CT93 C 3 3 Littoridina sphinctostoma 3 110CT93 C 3 3 Mediomastus ambiseta 3 110CT93 C 3 10 Streblospio benedicti 1 110CT93 C 3 10 Mediomastus ambiseta 2 110CT93 D 1 3 Rhynchocoel (unidentified) 2 110CT93 D 1 3 Streblospio benedicti 1 110CT93 D 1 3 Monoculodes sp. 1 110CT93 D 1 3 Mediomastus ambiseta 7 110CT93 D 1 10 Parandalia ocularis 2 110CT93 D 1 10 Mediomastus ambiseta 4 110CT93 D 2 3 Rhynchocoel (unidentified) 1 110CT93 D 2 3 Streblospio benedicti 3 110CT93 D 2 3 Littoridina sphinctostoma 4 110CT93 D 2 3 Mediomastus ambiseta 5 110CT93 D 2 10 Littoridina sphinctostoma 1 110CT93 D 2 10 Mediomastus ambiseta 2 110CT93 D 3 3 Rhynchocoel (unidentified) 1 110CT93 D 3 3 Streblospio benedicti 2 110CT93 D 3 3 Turbellaria (unidentified) 1 110CT93 D 3 3 Mediomastus ambiseta 5 110CT93 D 3 10 Littoridina sphinctostoma 1 110CT93 D 3 10 Mediomastus ambiseta 1 05JAN94 A 1 3 Streblospio benedicti 5 05JAN94 A 1 3 Mulinia lateralis 2 05JAN94 A 1 3 Littoridina sphinctostoma 17 05JAN94 A 1 3 Thompsonula sp. 4 05JAN94 A 1 3 Mediomastus ambiseta 3 05JAN94 A 1 10 Mediomastus ambiseta 1 05JAN94 A 2 3 Eteone heteropoda 1 05JAN94 A 2 3 Streblospio benedicti 7 05JAN94 A 2 3 Mulinia lateralis 4 05JAN94 A 2 3 Littoridina sphinctostoma 8 05JAN94 A 2 3 Mediomastus ambiseta 6 05JAN94 A 2 10 Neanthes succinea 1 05JAN94 A 2 10 Hobsonia florida 1 05JAN94 A 2 10 Mediomastus ambiseta 3 05JAN94 A 3 3 Streblospio benedicti 6 05JAN94 A 3 3 Mulinia lateralis 2 05JAN94 A 3 3 Chironomid larvae 1 05JAN94 A 3 3 Rangia cuneata 2 05JAN94 A 3 3 Littoridina sphinctostoma 7 05JAN94 A 3 3 Thompsonula sp. 1 05JAN94 A 3 10 Capitella capitata 2 05JAN94 A 3 10 Chironomid larvae 1 05JAN94 B 1 3 Streblospio benedicti 2 05JAN94 B 1 3 Capitella capitata 1 05JAN94 B 1 3 Mediomastus ambiseta 11 05JAN94 B 1 3 Glycinde nordmanni 1 05JAN94 B 1 10 Neanthes succinea 1 05JAN94 B 1 10 Parandalia ocularis 1 05JAN94 B 1 10 Mediomastus ambiseta 1 05JAN94 B 1 10 Glycinde nordmanni 1 05JAN94 B 2 3 Streblospio benedicti 8 05JAN94 B 2 3 Mulinia lateralis 2 05JAN94 B 2 3 Littoridina sphinctostoma 1 05JAN94 B 2 3 Thompsonula sp. 1 05JAN94 B 2 3 Mediomastus ambiseta 7 05JAN94 B 2 3 Glycinde nordmanni 1 05JAN94 B 2 10 Capitelia capitata 1 05JAN94 B 2 10 Glycinde nordmanni 1 05JAN94 B 3 3 Streblospio benedicti 3 05JAN94 B 3 3 Capitelia capitata 1 05JAN94 B 3 3 Chironomid larvae 1 05JAN94 B 3 3 Mediomastus ambiseta 15 05JAN94 B 3 10 Capitelia capitata 1 05JAN94 B 3 10 Parandalia ocularis 1 05JAN94 B 3 10 Mediomastus ambiseta 4 05JAN94 C 1 3 Rhynchocoel (unidentified) 3 05JAN94 C 1 3 Streblospio benedicti 2 05JAN94 C 1 3 Mulinia lateralis 1 05JAN94 C 1 3 Macoma mitchelli 1 05JAN94 C 1 3 Littoridina sphinctostoma 5 05JAN94 C 1 3 Mediomastus ambiseta 17 05JAN94 C 1 10 Mediomastus ambiseta 3 05JAN94 C 2 3 Rhynchocoel (unidentified) 2 05JAN94 C 2 3 Diopatra cuprea 1 05JAN94 C 2 3 Streblospio benedicti 5 05JAN94 C 2 3 Mulinia lateralis 3 05JAN94 C 2 3 Cyclaspis varians 2 05JAN94 C 2 3 Littoridina sphinctostoma 10 05JAN94 C 2 3 Thompsonula sp. 1 05JAN94 C 2 3 Mediomastus ambiseta 28 05JAN94 C 2 10 Littoridina sphinctostoma 1 05JAN94 C 2 10 Mediomastus ambiseta 1 05JAN94 C 3 3 Streblospio benedicti 11 05JAN94 C 3 3 Pectinaria gouldii 1 05JAN94 C 3 3 Mulinia lateralis 4 05JAN94 C 3 3 Littoridina sphinctostoma 9 05JAN94 C 3 3 Mediomastus ambiseta 31 05JAN94 C 3 10 Mediomastus ambiseta 1 05JAN94 D 1 3 Rhynchocoel (unidentified) 1 05JAN94 D 1 3 Streblospio benedicti 7 05JAN94 D 1 3 Paraprionospio pinnata 1 05JAN94 D 1 3 Spiochaetopterus costarum 2 05JAN94 D 1 3 Littoridina sphinctostoma 6 05JAN94 D 1 3 Mediomastus ambiseta 16 05JAN94 D 1 3 Glycinde nordmanni 1 05JAN94 D 1 10 Mediomastus ambiseta 2 05JAN94 D 2 3 Anthozoa (unidentified) 1 05JAN94 D 2 3 Rhynchocoel (unidentified) 2 05JAN94 D 2 3 Streblospio benedicti 5 05JAN94 D 2 3 Scolelepis texana 1 05JAN94 D 2 3 Spiochaetopterus costarum 1 05JAN94 D 2 3 Ensis minor 1 05JAN94 D 2 3 Ampelisca abdita 1 05JAN94 D 2 3 Parandalia ocularis 1 05JAN94 D 2 3 Mediomastus ambiseta 18 05JAN94 D 2 10 Spiochaetopterus costarum 1 05JAN94 D 2 10 Mediomastus ambiseta 3 05JAN94 D 3 3 Diopatra cuprea 1 05JAN94 D 3 3 Streblospio benedicti 3 05JAN94 D 3 3 Scolelepis texana 1 05JAN94 D 3 3 Molgula manhattensis 1 05JAN94 D 3 3 Mediomastus ambiseta 21 05JAN94 D 3 10 Spiochaetopterus costarum 3 05JAN94 D 3 10 Mediomastus ambiseta 1 07APR94 A 1 3 Streblospio benedicti 94 07APR94 A 1 3 Odostomia sp. 3 07APR94 A 1 3 Mulinia lateralis 5 07APR94 A 1 3 Monoculodes sp. 3 07APR94 A 1 3 Littoridina sphinctostoma 14 07APR94 A 1 3 Mediomastus ambiseta 2 07APR94 A 1 10 Streblospio benedicti 3 07APR94 A 1 10 Mediomastus ambiseta 3 07APR94 A 2 3 Streblospio benedicti 56 07APR94 A 2 3 Capitella capitata 1 07APR94 A 2 3 Mulinia lateralis 3 07APR94 A 2 3 Monoculodes sp. 3 07APR94 A 2 3 Littoridina sphinctostoma 22 07APR94 A 2 3 Mediomastus ambiseta 2 07APR94 A 2 10 Streblospio benedicti 1 07APR94 A 2 10 Mediomastus ambiseta 1 07APR94 A 3 3 Oligochaetes (unidentified) 2 07APR94 A 3 3 Streblospio benedicti 42 07APR94 A 3 3 Chironomid larvae 1 07APR94 A 3 3 Rangia cuneata 1 07APR94 A 3 3 Littoridina sphinctostoma 21 07APR94 A 3 10 Mediomastus ambiseta 2 07APR94 B 1 3 Streblospio benedicti 86 07APR94 B 1 3 Capitella capitata 3 07APR94 B 1 3 Mulinia lateralis 1 07APR94 B 1 3 Macoma mitchelli 1 07APR94 B 1 3 Littoridina sphinctostoma 8 07APR94 B 1 3 Mediomastus ambiseta 10 07APR94 B 1 10 Streblospio benedicti 1 07APR94 B 1 10 Capitella capitata 3 07APR94 B 1 10 Mediomastus ambiseta 5 07APR94 B 2 3 Streblospio benedicti 122 07APR94 B 2 3 Capitella capitata 2 07APR94 B 2 3 Littoridina sphinctostoma 4 07APR94 B 2 3 Mediomastus ambiseta 14 07APR94 B 2 10 Capitella capitata 1 07APR94 B 2 10 Mediomastus ambiseta 14 07APR94 B 3 3 Rhynchocoel (unidentified) 1 07APR94 B 3 3 Eteone heteropoda 1 07APR94 B 3 3 Streblospio benedicti 96 07APR94 B 3 3 Capitella capitata 2 07APR94 B 3 3 Cyclaspis varians 1 07APR94 B 3 3 Nudibranchia (unidentified) 1 07APR94 B 3 3 Littoridina sphinctostoma 4 07APR94 B 3 3 Mediomastus ambiseta 7 07APR94 B 3 10 Streblospio benedicti 1 07APR94 B 3 10 Capitella capitata 1 07APR94 B 3 10 Mediomastus ambiseta 1 07APR94 C 1 3 Streblospio benedicti 18 07APR94 C 1 3 Mulinia lateralis 4 07APR94 C 1 3 Cyclaspis varians 3 07APR94 C 1 3 Littoridina sphinctostoma 10 07APR94 C 1 3 Mediomastus ambiseta 33 07APR94 C 1 10 Streblospio benedicti 1 07APR94 C 1 10 Capitella capitata 1 07APR94 C 1 10 Mediomastus ambiseta 6 07APR94 C 1 10 Glycinde nordmanni 1 07APR94 C 2 3 Streblospio benedicti 19 07APR94 C 2 3 Mulinia lateralis 3 07APR94 C 2 3 Cyclaspis varians 2 07APR94 C 2 3 Monoculodes sp. 1 07APR94 C 2 3 Pyramidella crenulata 1 07APR94 C 2 3 Littoridina sphinctostoma 7 07APR94 C 2 3 Parandalia ocularis 3 07APR94 C 2 3 Mediomastus ambiseta 27 07APR94 C 2 10 Parandalia ocularis 1 07APR94 C 2 10 Mediomastus ambiseta 15 07APR94 C 3 3 Streblospio benedicti 12 07APR94 C 3 3 Cyclaspis vahans 2 07APR94 C 3 3 Littoridina sphinctostoma 13 07APR94 C 3 3 Diastylis sp. 1 07APR94 C 3 3 Mediomastus ambiseta 30 07APR94 C 3 10 Macoma mitchelli 1 07APR94 C 3 10 Mediomastus ambiseta 2 07APR94 D 1 3 Streblospio benedicti 2 07APR94 D 1 3 Mulinia lateralis 1 07APR94 D 1 3 Microprotopus spp. 1 07APR94 D 1 3 Oxyurostylis smithi 1 07APR94 D 1 3 Mediomastus ambiseta 16 07APR94 D 1 10 Spiochaetopterus costarum 1 07APR94 D 1 10 Haploscoloplos foliosus 1 07APR94 D 1 10 Mediomastus ambiseta 6 07APR94 D 2 3 Streblospio benedicti 1 07APR94 D 2 3 Spiochaetopterus costarum 2 07APR94 D 2 3 Parandalia ocularis 1 07APR94 D 2 3 Mediomastus ambiseta 16 07APR94 D 2 3 Glycinde nordmanni 2 07APR94 D 2 10 Mediomastus ambiseta 6 07APR94 D 3 3 Rhynchocoel (unidentified) 1 07APR94 D 3 3 Streblospio benedicti 3 07APR94 D 3 3 Paraprionospio pinnata 1 07APR94 D 3 3 Scolelepis texana 1 07APR94 D 3 3 Spiochaetopterus costarum 1 07APR94 D 3 3 Thompsonula sp. 1 07APR94 D 3 3 Parandalia ocularis 1 07APR94 D 3 3 Mediomastus ambiseta 43 07APR94 D 3 10 Spiochaetopterus costarum 1 07APR94 D 3 10 Parandalia ocularis 1 07APR94 D 3 10 Mediomastus ambiseta 4 NITROGEN LOSSES TO ESTUARINE SEDIMENTS INTRODUCTION A major goal of the Texas Water Development Board is to create nitrogen budgets for all Texas bays and estuaries. Nitrogen is the key element to controlling productivity and river inflow is a significant source of nitrogen. A simple budget would account for nitrogen entering the bay via freshwater inflow, how it is captured and transformed into biomass, and finally how it is lost to the system. One aspect of nitrogen loss is very poorly understood. How much nitrogen is buried and lost the system? Data on the nitrogen content of deep sediments can help answer this question. METHODS Several Estuaries have been studied. The Sabine-Neches and Trinity-San Jacinto Estuaries were sampled in 1993. The Lavaca-Colorado and Guadalupe Estuaries were sampled in 1990, and resampled in 1992. The Nueces Estuary and Baffin Bay were sampled in 1991. The station locations for the Sabine-Neches and Trinity-San Jacinto Estuaries are given in Table 1. The standard, long-term station locations were used in the Lavaca-Colorado, Guadalupe, and Nueces Estuaries and Baffin Bay. These locations are given in chapter 1. In addition, two other stations were sampled in Baffin Bay. These stations named for the Baffin Bay channel marker numbers at which the stations were taken. Our approach is to take sediments cores and measure nitrogen changes with respect to sediment depth. Cores are taken to a depth of 1 m. One-cm sediment sections are taken at the depth intervals listed. The sediment is dried, ground up, and homogenized. Carbon and nitrogen content, as a percent dry weight of sediment, is measured using a CHN analyzer. are RESULTS Nitrogen and carbon values for all measurements taken are shown in Tables 2-7. In recent sampling in the Sabine-Neches and Trinity-San Jacinto Estuaries sampling has been performed log-scale with respect to depth (Tables 2 and 3). Half the samples on a were concentrated in the upper 15 cm of sediment. In older sampling in the Lavaca- Colorado, Guadalupe, Nueces Estuaries and in Baffin Bay the samples were taken on a linear scale (Tables 4-7). All of these samples were taken at 10 cm depth intervals. The average for all samples at each depth interval is given in Table 8. In general, more N seems to be flowing into the northern estuaries, because content is higher in surface sediments of the Sabine-Neches and Trinity-San Jacinto Estuaries. One exception is Baffin Bay, where the nitrogen contents are about double This is any other system. very odd, and resampling during 1994 will be performed to confirm this trend. The N content at all stations sampled in the Sabine-Neches Estuary is average shown in Figure 1. Stations 1-6 represent a gradient from the head of the estuary to the sea. There N profiles at each station is similar. The N content at every depth average interval at all stations is shown in Figure 2. It appears that the labile zone extends to a depth of 40 cm. The refractory zone is not really constant until a depth of 80 cm. The average N content at all stations sampled in the Trinity-San Jacinto Estuary is shown in Figure 3. Stations 1-6 represent a gradient from the head of the estuary to the sea. There N profiles appear to be similar at all stations, except perhaps stations 3 and 4. At station 4, no trend is present. At station 3 the trend is inverse, that is it increases with depth. The average N content at every depth interval at all stations is shown in Figure 4. It that the labile zone extends to a depth of 15 cm. The appears refractory zone is constant from a depth of 20-100 cm. The average N content at all stations sampled in the Lavaca-Colorado Estuary is shown in Figure 5. Stations A-D represent a gradient from the head of the estuary to the sea. There N profiles appear to be similar at all stations. It appears nitrogen content decreases away from the head of the estuary. The average N content at every depth interval at all stations is shown in Figure 6. It appears that the labile zone extends to a depth of 40 cm. The refractory zone has an unusual pattern. It decreases to a depth of 60 cm, then increases to 100 cm. The average N content at all stations sampled in the Guadalupe Estuary is shown in Figure 7. Stations A-D represent a gradient from the head of the estuary to the sea. There N profiles to be similar at all stations. nitrogen content appear It appears decreases N content at away from the head of the estuary. The average every depth interval at all stations is shown in Figure 8. It appears that the labile zone extends to a depth of 50 cm. The Labile zone has an unusual pattern. It increases to a depth of 30 cm. The Refractory zone decreases to a depth of 70 cm, and is constant to 100 cm. The average N content at all stations sampled in the Nueces Estuary is shown in Figure 9. Stations A-E represent a gradient from the head of the estuary to the sea. There N profiles appear to be similar at all stations. It appears nitrogen content decreases away from the head of the estuary. An exception is that the highest values occur at Station C. The average N content at every depth interval at all stations is shown in Figure 10. It appears that the labile zone extends to a depth of 50 cm. The Labile zone decreases at a low rate. The Refractory zone decreases from a depth of SO­SO cm at a low rate, and then decreases dramatically to 100 cm. The average N content at all stations sampled in Baffin Bay is shown in Figure 11. Stations 24-6 represent a gradient from the head of the estuary to the sea. Station 24 is near the junction of the three secondary bays that empty into Baffin Bay. There N profiles appear to be similar at all stations. It appears nitrogen content decreases away from the head of the estuary. However, the values are very high, nearly double that found in other bays. The average N content at every depth interval at all stations is shown in Figure 12. It appears that the labile zone extends to a depth of 40 cm. The Refractory zone has an unusual pattern. It increases from a depth of 40 to 80 cm, then decreases to 100 cm. DISCUSSION If N comes into bays via the rivers and it is buried, then we would expect that there would be higher N values in the sediments at the head of estuaries. This is because the river empties into the bay, and more nitrogen could be trapped in the upper reaches of the bay. The station trends at all estuaries confirm this hypothesis (Figures 1,3, 5,7, 9, and 11). The only estuary where the trend is not strong is in the Sabine-Neches estuary. If N is utilized, or transformed in the biologically active labile zone, then there should be higher values of N in upper layers of sediment and lower values at lower layers in the refractory zone. This hypothesis is confirmed by the trends seen in the estuary-wide average N content (Figures 2,4, 6,8, 10, and 12). The labile zone appears to be as deep as 40 cm in most estuaries. The only exception is Trinity-San Jacinto, where the level is at about 20 cm. The finding that the refractory zone is as deep as 40 is surprising, but this cm due and difficult to know how much an area being sampled is subject to these disturbances. An alternative hypothesis to a labile and refractory zone is that there is simply N could be to anthropogenic influences, e.g., shrimping dredging. It is very more coming into bays today than at previous times. This would also explain the vertical distribution of N content in sediments. Man can influence another key component that affects N loss. In general, it is thought that the sedimentation rate in Texas estuaries is about 1 cm/ 100 years (Behrens, 1980). However, recent water projects, particularly dams, have probably decreased this rate. An average background level, i.e., the average content of N at about 40 cm is about 0.05% (Table 8). The surface N content is about 0.1%, so the change average is a factor of 2. This implies that half of the nitrogen arriving at the sediment surface is lost to the system via burial. REFERENCES Behrens, E.W. 1980. On sedimentation rates and porosity. Marine Geology Letters, 35.M11-Ml6. Table 1. Locations of stations sampled in 1993. A. Sabine-Neches Estuary. Station Latitude (N) Longitude (W) 1 29.57.’13.4" 93.49.’48.4" 2 29.54’.39.9" 93.48.’46.2" 3 29.52.’29.3" 93.48.’02.0" 4 29.51.’59.6" 93.51.’31.9" 5 29.49.’59.1" 93.51.’42.0" 6 29.47.’58.9" 93.55.’03.0" B. Trinity-San Jacinto Estuary. Station Latitude (N) Longitude (W) 1 29.42.’06.7" 94.44.’37.9" 2 29.37.'45.2" 94.49.’42.4" 3 29.33.’20.8" 94.59.’44.1" 4 29.22.’59.0" 94.50.’34.4" 5 29.26.’36.1" 94.43.’10.4" Table 2. Sabine-Neches Estuary sediment elemental composition. Depth in cm, Nitrogen and Carbon in % dry weight of sediment. Date Station Depth Nitrogen Carbon 050CT93 1 0 0.060 0.590 050CT93 1 2 0.086 0.938 050CT93 1 5 0.058 0.636 050CT93 1 10 0.038 0.391 050CT93 1 15 0.042 0.466 050CT93 1 20 0.037 0.391 050CT93 1 40 0.025 0.229 050CT93 1 60 0.022 0.200 050CT93 1 80 0.034 0.438 050CT93 1 100 0.033 0.392 050CT93 2 0 0.121 1.248 050CT93 2 2 0.090 0.921 050CT93 2 5 0.085 0.892 050CT93 2 10 0.071 0.771 050CT93 2 15 0.059 0.630 050CT93 2 20 0.115 1.283 050CT93 2 40 0.040 0.403 050CT93 2 60 0.043 0.480 050CT93 2 80 0.045 0.524 050CT93 2 100 0.049 0.578 050CT93 3 0 0.100 0.947 050CT93 3 2 0.065 0.677 050CT93 3 5 0.071 0.743 050CT93 3 10 0.075 0.793 050CT93 3 15 0.070 0.717 050CT93 3 20 0.103 1.196 050CT93 3 40 0.060 0.697 050CT93 3 60 0.050 0.642 050CT93 3 80 0.045 0.559 050CT93 3 100 0.050 0.594 050CT93 4 0 0.067 0.608 050CT93 4 2 0.054 0.501 050CT93 4 5 0.054 0.524 050CT93 4 10 0.062 0.631 050CT93 4 15 0.081 0.842 050CT93 4 20 0.052 0.506 050CT93 4 40 0.122 1.215 050CT93 4 60 0.094 1.034 050CT93 4 80 0.032 0.437 050CT93 4 100 0.045 0.764 050CT93 5 0 0.084 0.786 050CT93 5 2 0.068 0.661 050CT93 5 5 0.061 0.600 050CT93 5 10 0.044 0.561 050CT93 5 15 0.046 0.554 050CT93 5 20 0.056 0.588 050CT93 5 40 0.054 0.618 050CT93 5 60 0.052 0.661 050CT93 5 80 0.065 0.812 050CT93 5 100 0.053 0.659 050CT93 6 0 0.065 0.907 050CT93 6 2 0.070 0.909 050CT93 6 5 0.073 1.057 050CT93 6 10 0.095 1.222 050CT93 6 15 0.074 0.939 050CT93 6 20 0.088 1.034 050CT93 6 40 0.044 0.585 050CT93 6 60 0.051 0.860 050CT93 6 80 0.041 0.529 050CT93 6 100 0.044 0.570 Table 3. Trinity-San Jacinto Estuary sediment elemental composition. Depth in cm Nitrogen and Carbon in % dry weight of sediment. Date Station 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 1 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 2 050CT93 3 050CT93 3 050CT93 3 050CT93 3 050CT93 3 050CT93 3 050CT93 3 050CT93 3 050CT93 3 Depth 0 2 5 10 15 20 40 60 80 100 0 2 5 10 15 20 40 60 80 100 0 2 5 10 15 20 40 60 80 Nitrogen 0.112 0.115 0.099 0.102 0.102 0.088 0.090 0.096 0.088 0.067 0.157 0.138 0.133 0.127 0.120 0.095 0.095 0.111 0.117 0.100 0.144 0.120 0.112 0.109 0.090 0.094 0.139 0.161 0.153 Carbon 1.020 1.035 1.014 1.105 1.177 1.104 1.234 1.213 0.859 0.672 1.328 1.240 1.255 1.218 1.230 0.999 0.986 1.001 1.163 1.114 1.241 1.072 1.032 1.021 0.887 0.885 1.234 1.347 1.233 050CT93 3 100 0.162 1.320 050CT93 4 0 0.045 0.478 050CT93 4 2 0.048 0.476 050CT93 4 5 0.056 0.599 050CT93 4 10 0.035 0.415 050CT93 4 15 0.038 0.470 050CT93 4 20 0.039 0.424 050CT93 4 40 0.051 0.551 050CT93 4 60 0.061 0.619 050CT93 4 80 0.028 0.403 050CT93 4 100 0.045 0.570 050CT93 5 0 0.168 1.314 050CT93 5 2 0.173 1.374 050CT93 5 5 0.092 0.861 050CT93 5 10 0.051 0.526 050CT93 5 15 0.085 0.818 050CT93 5 20 0.082 0.797 050CT93 5 40 0.050 0.489 050CT93 5 60 0.039 0.446 050CT93 5 80 0.030 0.349 050CT93 5 100 0.031 0.430 050CT93 6 0 0.171 1.638 050CT93 6 2 0.081 1.314 050CT93 6 5 0.083 1.359 050CT93 6 10 0.075 0.844 050CT93 6 15 0.076 0.803 050CT93 6 20 0.080 0.841 050CT93 6 40 0.058 0.629 050CT93 6 60 0.067 0.702 050CT93 6 80 0.063 0.695 050CT93 6 100 0.061 0.735 Table 4. Lavaca-Colorado Estuary sediment elemental composition. Rep=replicate Depth in cm, Nitrogen and Carbon in % dry weight of sediment. Date Station Rep Depth Nitrogen Carbon 230CT90 A 1 10 0.067 0.875 230CT90 A 1 20 0.047 0.900 230CT90 A 1 30 0.033 1.151 230CT90 A 1 40 0.026 1.338 230CT90 A 2 10 0.057 0.873 230CT90 A 2 20 0.063 1.023 230CT90 A 2 30 0.049 0.957 230CT90 A 2 40 0.065 1.387 230CT90 A 2 50 0.048 1.068 230CT90 A 2 60 0.039 0.811 230CT90 A 2 70 0.037 0.852 230CT90 A 2 80 0.030 0.787 230CT90 A 2 90 0.026 0.656 230CT90 A 2 100 0.041 1.077 230CT90 B 1 20 0.072 1.266 230CT90 B 1 30 0.083 1.367 230CT90 B 1 40 0.096 1.664 230CT90 B 1 50 0.028 1.554 230CT90 B 1 60 0.011 1.206 230CT90 B 1 100 0.003 1.553 230CT90 B 2 10 0.068 1.234 230CT90 B 2 20 0.042 1.065 230CT90 B 2 30 0.061 1.522 230CT90 B 2 40 0.079 1.705 230CT90 B 2 50 0.069 1.522 230CT90 B 2 70 0.042 1.296 230CT90 B 2 80 0.039 0.937 230CT90 B 2 90 0.044 1.300 230CT90 B 2 100 0.064 1.654 230CT90 C 1 20 0.009 1.757 230CT90 C 1 40 0.048 1.545 230CT90 C 1 50 0.020 1.615 230CT90 C 1 60 0.020 1.496 230CT90 C 1 70 0.064 1.256 230CT90 C 1 80 0.053 1.331 230CT90 C 1 90 0.059 1.381 230CT90 C 1 100 0.068 1.521 230CT90 C 2 10 0.061 2.061 230CT90 C 2 20 0.052 2.789 230CT90 C 2 30 0.047 1.682 230CT90 C 2 40 0.042 1.333 230CT90 C 2 50 0.050 1.665 230CT90 C 2 60 0.038 1.303 230CT90 C 2 70 0.045 1.120 230CT90 C 2 80 0.045 1.252 230CT90 C 2 90 0.085 1.556 230CT90 C 2 100 0.046 1.491 230CT90 D 1 10 0.062 1.429 230CT90 D 1 20 0.005 0.377 230CT90 D 1 40 0.033 0.631 230CT90 D 1 50 0.026 0.569 230CT90 D 1 60 0.010 0.386 230CT90 D 1 70 0.054 1.205 230CT90 D 1 80 0.027 0.614 230CT90 D 1 90 0.034 0.881 230CT90 D 1 100 0.159 0.889 230CT90 D 2 10 0.057 1.254 230CT90 D 2 20 0.053 1.094 230CT90 D 2 30 0.041 0.832 230CT90 D 2 40 0.031 0.700 230CT90 D 2 50 0.023 0.521 230CT90 D 2 60 0.015 0.424 230CT90 D 2 70 0.007 0.350 230CT90 D 2 80 0.020 0.766 230CT90 D 2 90 0.028 0.910 230CT90 D 2 100 0.025 0.828 060CT92 A 1 10 0.104 1.489 060CT92 A 1 20 0.117 1.674 060CT92 A 1 30 0.065 1.233 060CT92 A 1 40 0.044 1.004 060CT92 A 1 50 0.044 1.174 060CT92 A 1 60 0.048 1.076 060CT92 A 1 70 0.061 1.388 060CT92 A 1 80 0.047 1.391 060CT92 A 1 90 0.049 1.103 060CT92 A 1 100 0.032 0.922 060CT92 B 1 10 0.070 1.282 060CT92 B 1 20 0.063 1.390 060CT92 B 1 30 0.063 1.386 060CT92 B 1 40 0.069 1.420 060CT92 B 1 50 0.076 1.453 060CT92 B 1 60 0.096 1.867 060CT92 B 1 70 0.069 1.434 060CT92 B 1 80 0.080 1.530 060CT92 B 1 90 0.078 1.558 060CT92 B 1 100 0.064 1.605 060CT92 C 1 10 0.053 1.412 060CT92 C 1 20 0.060 1.290 060CT92 C 1 30 0.044 1.545 060CT92 C 1 40 0.050 1.632 060CT92 C 1 50 0.053 1.618 060CT92 C 1 60 0.044 1.597 060CT92 C 1 70 0.044 2.073 060CT92 C 1 80 0.040 1.727 060CT92 C 1 90 0.037 1.521 060CT92 C 1 100 0.033 1.560 060CT92 D 1 10 0.052 1.386 060CT92 D 1 20 0.042 1.229 060CT92 D 1 30 0.028 0.663 060CT92 D 1 40 0.019 0.515 060CT92 D 1 50 0.019 0.587 060CT92 D 1 60 0.023 0.704 060CT92 D 1 70 0.032 0.951 060CT92 D 1 80 0.028 0.967 060CT92 D 1 90 0.032 1.076 060CT92 D 1 100 0.020 0.809 Table 5. Guadalupe Estuary sediment elemental composition. Rep=replicate, Depth in cm, Nitrogen and Carbon in % dry weight of sediment. Date Station Rep Depth Nitrogen Carbon 190CT90 A 1 10 0.060 3.403 190CT90 A 1 20 0.047 3.305 190CT90 A 1 30 0.059 3.459 190CT90 A 1 40 0.049 3.356 190CT90 A 1 50 0.036 4.028 190CT90 A 1 60 0.052 4.460 190CT90 A 1 70 0.041 4.146 190CT90 A 1 80 0.026 3.543 190CT90 A 1 90 0.028 3.544 190CT90 A 1 100 0.101 3.291 190CT90 A 2 10 0.089 3.564 190CT90 A 2 20 0.084 3.704 190CT90 A 2 30 0.090 3.910 190CT90 A 2 40 0.077 3.755 190CT90 A 2 50 0.050 3.803 190CT90 A 2 60 0.053 3.932 190CT90 A 2 70 0.044 4.288 190CT90 A 2 80 0.054 3.847 190CT90 A 2 90 0.054 3.652 190CT90 A 2 100 0.066 2.729 190CT90 B 1 10 0.084 3.615 190CT90 B 1 20 0.073 3.620 190CT90 B 1 30 0.073 3.489 190CT90 B 1 40 0.071 3.461 190CT90 B 1 50 0.058 3.908 190CT90 B 1 60 0.060 3.903 190CT90 B 1 70 0.059 3.846 190CT90 B 1 80 0.062 2.309 190CT90 B 1 90 0.042 2.579 190CT90 B 1 100 0.059 3.142 190CT90 B 2 20 0.075 3.626 190CT90 B 2 30 0.122 3.510 190CT90 B 2 40 0.078 3.694 190CT90 B 2 50 0.073 4.106 190CT90 B 2 60 0.061 3.132 190CT90 B 2 70 0.071 3.340 190CT90 B 2 80 0.057 3.329 190CT90 B 2 90 0.069 3.354 190CT90 B 2 100 0.058 3.089 190CT90 C 1 10 0.037 2.823 190CT90 C 1 20 0.076 2.571 190CT90 C 1 30 0.045 2.792 190CT90 C 1 40 0.052 2.648 190CT90 C 1 50 0.036 2.975 190CT90 C 1 60 0.069 3.121 190CT90 C 1 70 0.038 2.736 190CT90 C 1 80 0.046 2.987 190CT90 C 1 90 0.027 3.132 190CT90 C 1 100 0.025 4.057 190CT90 C 2 10 0.022 2.566 190CT90 C 2 20 0.012 2.719 190CT90 C 2 30 0.065 3.152 190CT90 C 2 40 0.047 3.057 190CT90 C 2 50 0.056 2.721 190CT90 C 2 60 0.051 3.131 190CT90 C 2 70 0.048 2.751 190CT90 C 2 80 0.025 3.205 190CT90 C 2 90 0.031 3.002 190CT90 C 2 100 0.011 3.284 190CT90 D 1 30 0.018 0.648 190CT90 D 1 100 0.004 3.674 190CT90 D 2 10 0.019 0.774 190CT90 D 2 20 0.022 0.921 190CT90 D 2 30 0.020 0.828 190CT90 D 2 50 0.026 0.916 190CT90 D 2 60 0.019 0.868 190CT90 D 2 70 0.016 0.789 190CT90 D 2 100 0.020 0.894 070CT92 A 1 10 0.076 3.902 070CT92 A 1 20 0.075 3.541 070CT92 A 1 30 0.082 3.566 070CT92 A 1 40 0.077 3.523 070CT92 A 1 50 0.074 4.427 070CT92 A 1 60 0.057 3.802 070CT92 A 1 70 0.058 3.692 070CT92 A 1 80 0.062 2.431 070CT92 A 1 90 0.062 3.140 070CT92 A 1 100 0.065 3.624 070CT92 B 1 10 0.091 4.292 070CT92 B 1 20 0.080 3.722 070CT92 B 1 30 0.086 3.708 070CT92 B 1 40 0.088 3.445 070CT92 B 1 50 0.086 3.673 070CT92 B 1 60 0.089 3.384 070CT92 B 1 70 0.091 3.398 070CT92 B 1 80 0.081 3.465 070CT92 B 1 90 0.087 3.187 070CT92 B 1 100 0.086 3.412 070CT92 C 1 10 0.065 2.690 070CT92 C 1 20 0.079 2.854 070CT92 C 1 30 0.054 2.731 070CT92 C 1 40 0.052 2.947 070CT92 C 1 50 0.075 2.875 070CT92 C 1 60 0.052 3.619 070CT92 C 1 70 0.046 3.120 070CT92 C 1 80 0.060 2.927 070CT92 C 1 90 0.064 3.117 070CT92 C 1 100 0.067 2.803 070CT92 D 1 10 0.030 0.936 070CT92 D 1 20 0.044 1.411 070CT92 D 1 30 0.055 1.785 070CT92 D 1 40 0.040 1.249 070CT92 D 1 50 0.040 1.200 . 070CT92 D 1 60 0.033 1.006 070CT92 D 1 70 0.040 1.253 070CT92 D 1 80 0.040 1.329 070CT92 D 1 90 0.024 1.067 070CT92 D 1 100 0.027 1.473 Table 6. Nueces Estuary sediment elemental composition. Rep=replicate, Depth in cm Nitrogen and Carbon in % dry weight of sediment. Date Station Rep Depth Nitrogen Carbon 160CT91 A 1 10 0.055 2.112 160CT91 A 1 20 0.050 2.036 160CT91 A 1 30 0.050 2.243 160CT91 A 1 40 0.055 2.354 160CT91 A 1 50 0.053 2.324 160CT91 A 1 60 0.054 2.339 160CT91 A 1 70 0.063 2.071 160CT91 A 1 80 0.060 2.288 160CT91 A 1 90 0.064 2.167 160CT91 A 1 100 0.057 2.131 160CT91 B 1 10 0.034 0.738 160CT91 B 1 20 0.029 0.930 160CT91 B 1 30 0.033 0.870 160CT91 B 1 40 0.031 0.991 160CT91 B 1 50 0.027 0.846 160CT91 B 1 60 0.032 0.918 160CT91 B 1 70 0.034 0.913 160CT91 B 1 80 0.038 0.967 160CT91 B 1 90 0.033 0.947 160CT91 B 1 100 0.029 0.784 160CT91 C 1 10 0.088 2.378 160CT91 C 1 20 0.087 2.436 160CT91 C 1 30 0.099 1.889 160CT91 C 1 40 0.111 2.258 160CT91 C 1 50 0.100 2.956 160CT91 C 1 60 0.089 2.048 160CT91 C 1 70 0.089 1.889 160CT91 C 1 80 0.089 1.827 160CT91 E 1 10 0.056 0.943 160CT91 E 1 20 0.052 0.804 160CT91 E 160CT91 E 160CT91 E 160CT91 E 160CT91 E 160CT91 E 160CT91 E 1 1 1 1 1 1 1 30 40 50 60 70 80 90 0.043 0.698 0.033 0.588 0.027 0.679 0.025 0.489 0.024 0.603 0.024 0.694 0.023 0.641 Table 7. Baffin Bay Estuary sediment elemental composition. Rep=replicate, Depth in cm, Nitrogen and Carbon in % dry weight of sediment. Date Station Rep Depth Nitrogen Carbon 170CT91 MKI2 1 10 0.175 1.530 170CT91 MKI2 1 20 0.150 1.592 170CT91 MKI2 1 30 0.155 0.455 170CT91 MKI2 1 40 0.134 1.364 170CT91 MKI2 1 50 0.147 2.888 170CT91 MKI2 1 60 0.208 4.211 170CT91 MKI2 1 70 0.211 3.880 170CT91 MKI2 1 80 0.222 4.236 170CT91 MKI2 1 90 0.172 3.645 170CT91 MKI2 1 100 0.168 2.505 170CT91 MKIB 1 10 0.201 1.628 170CT91 MKIB 1 20 0.151 1.793 170CT91 MKIB 1 30 0.143 1.338 170CT91 MKIB 1 40 0.155 1.435 170CT91 MKIB 1 50 0.146 1.386 170CT91 MKIB 1 60 0.167 1.901 170CT91 MKIB 1 70 0.145 2.292 170CT91 MKIB 1 80 0.186 3.549 170CT91 MKIB 1 90 0.187 2.943 170CT91 MKIB 1 100 0.181 3.376 170CT91 MK24 1 10 0.155 1.661 170CT91 MK24 1 20 0.135 1.376 170CT91 MK24 1 30 0.123 1.260 170CT91 MK24 1 40 0.138 1.425 170CT91 MK24 1 50 0.166 2.114 170CT91 MK24 1 60 0.119 1.718 170CT91 MK24 1 70 0.212 3.751 170CT91 MK24 1 80 0.170 4.632 170CT91 MK24 1 90 0.152 3.580 170CT91 MK24 1 100 0.148 2.426 170CT91 MK6 1 10 0.233 1.918 170CT91 MK6 1 20 0.151 1.496 170CT91 MK6 1 30 0.131 1.477 170CT91 MK6 1 40 0.103 1.466 170CT91 MK6 1 50 0.179 2.766 170CT91 MK6 1 60 0.195 4.651 170CT91 MK6 1 70 0.163 2.325 170CT91 MK6 1 80 0.221 3.516 170CT91 MK6 1 90 0.207 3.627 170CT91 MK6 1 100 0.202 3.021 Table 8. Average vertical distribution of N content (%) at all stations all estuaries among studied. A. Northern estuaries. SN=Sabine-Neches and TJ=Trinity-San Jacinto. B. Southern estuaries. LC=Lavaca-Colorado, GE=Guadalupe, NC=Nueces, and BB=Baffin Bay. A. Depth SN TJ 0 0.083 0.133 2 0.072 0.113 5 0.067 0.096 10 0.064 0.083 15 0.062 0.085 20 0.075 0.080 40 0.058 0.081 60 0.052 0.089 80 0.044 0.080 100 0.046 0.078 B. Depth LC GE NC BB 10 0.065 0.057 0.058 0.191 20 0.052 0.061 0.055 0.147 30 0.051 0.064 0.056 0.138 40 0.050 0.063 0.058 0.133 50 0.041 0.055 0.052 0.160 60 0.034 0.054 0.050 0.172 70 0.046 0.050 0.053 0.183 80 0.041 0.051 0.053 0.200 90 0.047 0.049 0.040 0.180 100 0.050 0.049 0.043 0.175 Fimirp 9 \/prtir.pl Hic:frihi itinn nf Mitrnnpn in fhp Fiphinp-Nprhpc Fq+iipr/ 141 Fiq. 3. Vertical distribution of Nitrogen at stations in the Trinity-San Jacinto Estuary. 142 Fig. 4. Vertical distribution of Nitrogen in the Trinity-San Jacinto Estuary. 143 144 145 146 147 148 Fig. 10. Vertical distribution of Nitrogen in the Nueces Estuary. 149 Fig. 11. Vertical distribution of Nitrogen at stations in the Baffin Bay. Fig. 12. Vertical distribution of Nitrogen in the Baffin Bay. 151