BULLETIN 25 Texas Memorial Museum Stratigraphic Occurrence and Correlation of Early Tertiary Vertebrate Faunas Trans-Pecos Texas Part I: Vieja Area By John Andrew Wilson AcceptedforPublication December5, 1977 THE TEXAS MEMORIAL MUSEUM 2400 Trinity, Austin, Texas 78705 CONTENTS ABSTRACT 1 INTRODUCTION 1 Acknowledgments 3 Previous Paleontological Work 6 Regional Geologic Setting 6 Laramide Structures 7 Volcanic Activity 7 Basin and Range Faulting 7 VIEJA AREA 7 Geologic Setting 7 Biostratigraphy Vieja Group 10 Jeff Conglomerate 10 Gill Breccia 11 Colmena Formation 11 Candelaria Local Fauna 13 Buckshot Ignimbrite 14 Chambers Tuff 14 Porvenir Local Fauna 18 Little Egypt Local Fauna 18 Capote Mountain Tuff 18 Vieja Group Undifferentiated 22 Ash Spring Local Fauna 23 Garren Group 23 Prietos Formation 23 Rancho Gaitan Local Fauna 23 Chronostratigraphy, Vieja Group 23 Radiochronology 23 Biochronology 24 Candelaria Local Fauna 24 Porvenir Local Fauna 25 Little Egypt Local Fauna 30 Airstrip Local Fauna 30 31 Rancho Gaitan Local Fauna 31 Paleoenvironment Chronostratigraphy and the Chadronian-Uintan Boundary 33 SUMMARY AND CONCLUSIONS 38 REFERENCES 39 III ILLUSTRATIONS Figure Page 1. Index map of Texas showing location of principle areas 2 2. Sketch map of volcanic centers and geologic quadrangles, Trans-Pecos Texas 4 3. Generalized stratigraphic sections with approximate position of vertebrate faunas and radiometric dated samples 5 4. Index to Vieja area 8 5. Index to Vieja stratigraphic units 9 6. Colmena section at mouth of Capote Creek 12 7. Chambers section at Big Cliff 15 8. Chambers section near Adobe Springs 16 9. Chambers section at Reeves bonebed 19 IV TABLES Table Page 1. Candelaria 1.f., late Uintan 13 2. Distribution of Candelaria l.f. in Colmena Tuff 13 3. Porvenir 1.f., early Chadronian 20 4. Taxa found in lower 100 ft. or beneath lower marker bed, Porvenir l.f. or "Big Red horizon" of Patterson 20 5. Taxa found at "red mound," Porvenir l.f. 21 6. Taxa found between o—BB ft. above lower marker bed, Porvenir l.f. or "Blue Cliff horizon" of Patterson 21 7. Taxa found at Adobe Springs area 21 8. Little Egypt 1.f., early Chadronian 21 9. Taxa found at Reeves bonebed, Little Egypt l.f 22 10. Taxa found at Chalk Gap Draw, Little Egypt l.f 22 11. Airstrip 1.f., Chadronian 22 12. Ash Spring 1.f., Chadronian 23 13. Rancho Gaitan l.f 24 14. Stratigraphic occurrence of currently identified taxa 27 ff. 15. Distribution of potassium-argon dates 30 16. Number of specimens of medium sized herbivores, Porvenir l.f 31 V STRATIGRAPHIC OCCURRENCE AND CORRELATION OF EARLY TERTIARY VERTEBRATE FAUNAS TRANS-PECOS TEXAS Part I. Vieja Area By John Andrew Wilson* ABSTRACT Stratigraphic positions for the Candelaria local fauna (late Uintan), the Porvenir local fauna (early Chadronian), Little Egypt, and Airstrip local faunas (Chadronian) within the Vieja Group are given. The Porvenir local fauna contains 32 genera in common with Chadronian faunas to the north and six generain common with the Lapoint fauna. K-AR dates of approximately 38 and 36 million years are below and above the Porvenir local fauna. A mammalian assemblage sufficientlydistinct to characterize a Duchesnean age is not present in the Vieja area. It is urged that the first appearance of Mesohippus mark the beginning of the Chadronian "age." INTRODUCTION The first Oligocene vertebrate fossil fauna from Trans-Pecos Texas was described byStovall (1948). Since that time vertebrate faunas from Paleocene, Eocene, Oligocene, and Miocene have been collected and most elements of the faunas have been described or are in press. This report is intended to bring together the results of this work and to correlate the Texas faunas from the Vieja area with those of other parts of North America. This biochronologic correlation is supplemented by radiochronological correlation where possible. Three general areas within Trans-Pecos Texas have produced the vertebrate faunas (fig. 1). From northwest to southeast these are (1) Sierra Vieja or Rim Rock Country, (2) a critical area including from south to ‘Research Associate, Texas Memorial Museum, Austin; Professor Emeritus, Department of Geological Sciences, The University of Texas at Austin. north, the Agua Fria Ranch, Bandera Mesa and Coffee Cup Ranch area, and (3) Big Bend National Park. The Tertiary sedimentary rocks of the BigBend area of west Texas were the last in the state to be divided into series whereas the subdivisions of the fossiliferous Tertiary of the easily accessible Coastal Plain were known by the turn of this century. The fossiliferous continental Pliocene deposits of the Panhandle were known by the 1890 s (Cope1893) and the type faunas for the Clarendonian and Hemphillian North American land-mammal ages were established by the 19305. On the other hand, in The Geologyof Texas volume 1, Stratigraphy (Sellards et al. 1933, p. 805), known as "3232," and the last summary publication on the stratigraphyof Texas, only one vertebrate fossil is mentioned as coming from Tertiary rocks of Trans-Pecos Texas. The specimen, TMM 41256-4, is a tooth of Hyracodon, an Oligocene rhinoceros, collected by C. L. Baker and identified by R. A. Stirton. At the presenttime, using approximate figures, there are known 28 genera of Paleocene mammals, 41 genera of Eocene mammals, 55 genera of Oligocene mammals, and 15 genera of Miocene mammals. In addition, there is a sizable herpetofauna, an invertebrate fauna, and floras that have not been studied. Except for two collections, one at the University of Oklahoma and the other at the Field Museum of Natural History, both of which are from the OligoceneChambers Tuff of the Vieja Group, the material is all at the Vertebrate PaleontologyLaboratory of the Texas Memorial Museum, The University of Texas at Austin. Vertebrate fossils were collected in the early 1950 s in the Tertiary of Big Bend National Park in connection with the pub 1 lication of the geologic map and bulletin (Maxwell et al. 1967) on the geology of the park. J. H. Quinn and I made small collections which were described in that publication (Wilson 1967) in order to document the presence of Paleocene and Eocene sediments. In the 1960 s my attention was directed to the the Vieja area where I worked in conjunctionwith the mapping program carried on by Professor R. K. DeFord. By the 1970 s the Vieja country had become less productive and a new area on the Agua Fria Quadrangle was discovered. The collections from each of the areas have been treated differently. The Vieja collection was divided into major taxonomic groups and distributed to experts for study and description. For example, rodents from the ViejaGroup were described by Albert E. Wood, horses by Paul 0. McGrew. This followed the pattern set by Professor Bryan Patterson, who made a major collection from the Chambers Tuff of the Vieja Group for the Field Museum of Natural History. In contrast the Paleocene fauna of the Black Peaks Formation of Big Bend National Park was treated as a single unit and described by Schiebout (1974). There are, however, parts of all faunas awaiting study. In addition, there are segments of the stratigraphic section that offer promise of successful collecting in the future. Volcaniclastic rocks of Trans-Pecos Texas (figs. 2,3) range from middle Eocene to Mio cene and have furnished large and important collections of vertebrate fossils. In addition, the volcanic rocks provide a framework of marker-beds which serve to subdivide locally thick accumulations of sediments. The occur rence together of the fossil faunas and vol canic rocks suitable for radiometric dating makes the area of study extremely important. Igneous dikes intrude the fossiliferous sedi ment and are suitable for radiometric dating. They furnish by their cross-cutting relation , ship, minimum age of the sediment. Radiometric dates pertinent to this study are taken from several sources and are given in table 15. Fig. 1 -Map of Texas with Presidio and Brewster counties shaded. Enlarged map shows location of principle fossil iferous areas. Big Bend National Park shaded. The late Cretaceous and early Tertiary sections of Trans-Pecos Texas encompass the Laramide interval. The Tornillo Basin stratigraphic section in Big Bend National Park, as given by Maxwell et al. (1967), and the sequence of events given by Wilson (1972) are similar to that given byTweeto (1975) for the southern Rocky Mountains. The Tornillo Basin was a structural basin during the Laramide interval like the Raton and Denver Basins to the north (Tweeto 1975, fig. 6). The Trans-Pecos area is technically part of the southern Rocky Mountains, but has been complicated by later volcanic activity and Basin and Range faulting. This report is divided into two parts. Part I deals with the Sierra Vieja area; Part II deals with the Agua Fria area and will be publishedlater. The purpose is to bring together stratigraphic and paleontologic information that is scattered in various publications and to present a correlation of the west Texas faunas with those elsewhere. Nothing significantly new can be added to the early Tertiary vertebrate faunas of Big Bend National Park at the present time. Acknowledgments The major financial support over the past25 years came from the University Research Institute and the Geology Foundation of The University of Texas at Austin. The National Science Foundation supported the work, in part, during the period from 1960-1965 through grants G 13270 and GP 1058. A grant, NGS 1439 (1975), was received from the National Geographic Society to make molds oftrackways. It is impossible to acknowledge individuallyall the people who have assisted me in this project. I have benefited from discussions with my colleagues on the faculty at The University of Texas at Austin, S. E. Clabaugh, R. K. DeFord, W. Langston, Jr., E. L. Lundelius, Jr., F. W. McDowell, and W. R. Muehlberger. The Albert Chambers and the Boyd Chambers families and Sheriff and Mrs. Francis Rooney were especially hospitable during our 3 Fig. 2.—Sketch map of Trans-Pecos Texas and adjacent Elms 1949); BM, Bofecillos Mountains QuadrangleMexico showing extent of extrusive igneous volcaniclastic (McKnight 1970); CM, Cathedral Mountain Quadranglerocks, known and inferred volcanic centers, and the loca-(McAnulty 1955); JG, Jordan Gap Quadrangle (Seward tion of geologic quadrangles. AF, Agua Fria Quadrangle 1959); TM, Tascotal Mesa Quadrangle (Erickson 1953). (Moon 1953); BH, Buck Hill Quadrangle (Goldich and Modified from Gorski (1970). 4 Devils County. Hudspeth HU, County; Culberson CULB, Park. NationalFormation Bend Graveyard Bi9 part in Compiled report. this of PartII in described be will field volcanic Texas Trans-Pecos the of sections stratigraphic 3-Generalized Fig and scale to not are sections Wilson, A. J. by modified and ) 1971 < Parker is F - D lithology V b diagrammatic. fossil dated of distribution the showing Mexico Chihuahua, Prietos, Cerros and composite ais 5Sectionsamples. dated potassium-argon and collections vertebrate in both Castolon, near is 6 section Mountains, Chisos and Flat Tornillo of section field work in the Vieja area. To other ranchers who generously gave me permission to enter their property, I am grateful. am especially grateful to Margaret and James B. Stevens for their unfailing physicaland scientific help in the field and in the laboratory. Anthony W. Walton studied the sedimentary rocks of the Vieja area. I thank him for permission to use information from his dissertation. I have also drawn from the theses and dissertations of W. B. Anderson, I. Ferrusquia- Villafranca, G. H. Heiken, C. J. Mankin, J. F. McKnight, J. T. Schulenberg, and P. C. Twiss. Judith A. Schiebout furnished me information on the tapirs and rhinos she is currentlystudying, and similarly Eric Gustafson furnished information on the carnivoresthat are the subject of his dissertation. Albert E. Wood has been consulted over the years concerning the rodents in the various faunas and their use in correlation. I am very grateful to him. Other collaborators have been Craig C. Black, Mary R. Dawson, Ann Forsten, John M, Harris, H. 0. Hofer, Paul 0. McGrew, Michael J. Novacek, and Fred Szalay. I am grateful to Craig C. Black, Mary R. Dawson, and Robert J. Emry for their constructive criticism of the manuscript. The conclusions, however, are my own. Abbreviations CRF Cretaceous rock fragments FMNH Field Museum of Natural History l. local fauna m. million yearsOU University of Oklahoma, Norman TMM Texas Memorial Museum, Austin VRF volcanic rock fragments Previous Vertebrate Paleontological Work A field party from the University of Oklahoma consisting of W.N. MacAnulty, D. E. Savage, and Wann Langston made the first collection of fossil vertebrates in the Vieja area in 1938; in 1940 Stovall and Savage returned. The collection was described by Stovall (1948). A second and larger collection was made in 1946 by Bryan Patterson and James H. Quinn for the Field Museum of Natural History. To my knowledge these are the only two organized field parties that had the express intent of searching for Tertiary vertebrate fossils in the Vieja area prior to 1950. Regional GeologicSetting The semi-arid climate and widespread outcrops make Trans-Pecos Texas one of the most interesting areas for geology and paleontology. A stratigraphic section extends from Precambrian to Pleistocene. The section has been folded and faulted, however, during a long and complex history. Appalachian Mountain type of folding, Rocky Mountain or Laramide folding and Basin and Range fault ing all intersect in the Big Bend area. In addition, during the late Eocene and Oligocenethe area was an active volcanic field. The youngest Cretaceous and the oldest Tertiary continental sediments are preservedin a structural low on Tornillo Flat in BigBend National Park. These sediments are debris that eroded from the eastern part of the area affected by the Laramide Orogeny. Paleocene fluvial and flood plain deposits lie on late Cretaceous sediment of similar origin. The Cretaceous sediments contain a reptilianfauna now attributed to Maestrichtian Age(Lawson 1976). The nearest similar stratigraphic sections to the north are found in Huerfano Park and the Denver Basin in Colorado, and the San Juan Basin, New Mexico. These more northerly basins, however, have not been so severely deformed as the Tornillo Basin. To the west, Morris (1967, 1968) has reported late Cretaceous and Paleocene fossil- bearing sediments on the peninsula of BajaCalifornia. To the southeast, the Rio Grande Embayment, and to the south the Sabinas Coal Basin and the Parras Basin all contain marine Paleocene sediments (Wilson 1972). In the Vieja area Uintan (late Eocene) and Chadronian (early Oligocene) tuffaceous sediments overlie late Cretaceous continental dinosaur-bearing rocks of the Picacho Fm. (Wolleben 1966). In the Agua Fria-Buck Hill 6 area early Uintan (early late Eocene) sediments lie on the Aguja Formation (Campanian (Moon 1953) and Boquillas Formation (Turonian-Santonian). Laramide Structures.-In the Big Bend area, structures that are commonly dated as havingbeen formed during the Laramide Orogeny are broad open anticlines and synclines and thrust faults. The Del Norte and SantiagoMountains to the north and northeast of the Tornillo Basin are Laramide structures (Maxwell et al. 1967). Cow Heaven anticline and Mariscal Mt. to the southwest and south are also Laramide structures. Other Laramide folds are found across the Rio Grande in Coahuila (Smith 1970) and in Chihuahua (Gries and Haenggi 1972). Erosion of the neighboring folds as they rose must have furnished the alluvial material that was trapped in the Tornillo Basin. Paleocene and early Eocene sediments and vertebrates have been found in a small area in the Big Bend National Park. Three localities within the park are located by Schiebout (1974, fig. 1). When the northern part of Coahuila and Chihuahua are more thoroughly explored, early Tertiary sediments may be found there also. Although Maxwell et al. (1967, p. 301) maintain that the lowest Tertiary rocks had a local origin, Schiebout (1974, p. 8) is of the opinion that some of the sediments are not locally derived, but come instead from the west or northwest. This would certainly be in accord with the general stratigraphic re lationships as outlined by Maxwell et al. (1967, p. 96). Volcanic Activity—The Big Bend area was the site of both intrusive and extrusive igneous activity. The oldest igneous rocks apparently are sills intruded into the late Cretaceous Aguja and Javelina Formations prior to the time of Laramide folding (Maxwell et al. 1967). Baumgarten (pers. comm. 1975) has traced one of the sills that is conformable with the folds at Maricsal Mt. and Cow Heaven anticline. The earliest extrusive igneousrock so far known is the unnamed basalt in ex- the Canoe Formation (middle Eocene) posed in the Crusher Section 1 and 2 (Maxwell et al. 1967, PI. IX) in Big Bend National Park. The climax of the igneous activity was during the early Oligocene when flow rock and ignimbrites accumulated over almost the entire area from the northern Davis Mountains to the Chisos Mountains in Big Bend National Park. By the Miocene most of the extrusive activity had ceased. An exception is the small volcano in the northern Vieja area described by DeFord and Bridges (1959). Dike swarms, however, continued to intrude during the Miocene (Dash et al. 1969). The interbedded fossiliferous sediments and volcanic flow rocks give the opportunity to correlate paleontological and radiometric dates from middle Eocene to Miocene. These are described for the separate areas. Basin and Range Fauiting—The Big Bend area lies on the eastern edge_ of the Basin and Range Province. The Rim Rock and associat ed faults in the Vieja area (DeFord 1958) cut Chadronian sediments and are attributed to Basin and Range faulting. In Big Bend National Park the western edge of the Sunken Block of Udden (1907) is bounded by the TerlinguaAbaja fault zone. Some of the faults within this zone cut Arikareean (early Miocene) sediments (Stevens 1969; Stevens et al. 1969). VIEJA AREA Geologic Setting The Sierra Vieja (figs. 1,4) is a north- northwest trending range that separates the high Davis Mountains and the Marfa Plateau from the rough area that borders the Rio Grande. Although it is only 10 to 15 miles from the rim of the Sierra Vieja (also called the Rim Rock), to the Rio Grande, the elevation drops about 2,000 feet. The Vieja area parallels the international border for about 60 miles from the village of Candelaria on the south to the Presidio-Jeff Davis County line on the north. The northern part of the area is drained by Dieciocho (Van Horn) and Quinn creeks, the southern part by CapoteCreek. The fossiliferous areas along these 7 Fig. 4.—lndex map to sections in the Vieja area. A. Section of Colmena Tuff at mouth of CapoteCreek, fig. 6. B. Section of Chambers Tuff at Big Cliff, fig. 7. C. Section of Chambers Tuff near Adobe Spring, fig. 8. D. Section of Chambers Tuff at Reeves Bonebed, fig. 9. E. Type section of Colmena Tuff (DeFord 1958). F. Type section of Chambers Tuff (DeFord 1958). G. Type section of Capote Mt. Tuff (DeFord 1958). H. Ash Spring locality (Harris 1967a, 1967b), Harris and Wood (1969). I. Rancho Gaitan locality (Ferrusquia-V. 1969). Fig. s.—lndex to stratigraphic units, potassium-argon dates, and mammalian local faunas in the SierraVieja or Rim Rock intermittent streams and their tributaries will be referred to as northern and southern areas. The Vieja area lies in the western part of the Davis Mountain volcanic field. Several centers of volcanism are known and others are inferred (fig. 2), but just which center contributed which particular extrusive igne rock to the Vieja area is unknown. An ous index to the stratigrpaphic succession of the flow rocks and intervening tuffs and tuffaceous sediments in the Vieja area is given in figure 5, together with the potassium-argon country. Modified from Walton (1972). The Ford Ranch bed is found only in the southern part of the Vieja area and is not close to any vertebrate localities. dates and the approximate position of the local faunas. Volcanic activity began in the Vieja area in the late Eocene, reached a peakduring the early Oligocene, and continued at least to the Miocene (Dash et al. 1969). The rim of the Sierra Vieja is formed bythe major scarp of the Rim Rock Fault zone, at the eastern edge of the Basin and RangePhysiographic Province. Displacement alongthe Rim Rock fault varies from 900 to 3,000feet (274-914m) (Baker 1934). The down- thrown or western block is cut by many lesser faults that tilt the sediments and flow rocks. 9 The faulting repeats the sequence in a verycomplicated manner. The igneous flow rocks are the most persistent marker beds and form the basis for mapping. In addition, other stratigraphic units, particularly widespreadtuffs, have been found that are reliable marker beds within the Chambers Tuff. Fortunately, the undisturbed stratigraphic section exposed along the fault scarp of the Rim Rock fault can be used to verify the superpositionof sediments and volcanic flows. A summaryof the geologic mapping and stratigraphic section is found in Twiss (1972). Biostratigraphy Vieja Group The lithologic units of the Vieja Group as described by DeFord (1958) form the framework of this section of the report. Vertebrate fossils have been collected at various places from some of the lithologic units. Where the collections were considered to be close enough stratigraphically and geographically they were grouped into local faunas. The vertical and horizontal extensions of the volume of rock partly occupied by the local fauna does not coincide with and is not to be interpreted as coincident with the lithologicunit. For example, the Colmena Tuff is a lithologic unit that has been divided by Walton (1972) into a northern conglomeratic facies and a southern volcanic rock facies. The southern facies in turn can be subdivided into a lower sorted facies, a middle lahar facies, and an upper glass facies. Fossil vertebrates have been found only in the southern area and most come from the sorted facies. One tooth was found in the middle or lahar facies and in the upper glass facies enough taxa were found in common with the lower facies so that all were grouped together in the Candelaria l.f. A brief lithologic description of each unit or marker bed within the Vieja Group is given for purposes of ease of field identification, without implying that the lithologic unit is isochronous, unless so stated. Attempts were made to relate vertebrate occurrences to useful marker beds. Three such beds occur in the Chambers Tuff but none in the Capote Mountain Tuff. The lithologic units up through the Chambers Tuff were deposited under varying conditions, including high energy. Thicknesses vary so much that the recording of footage was felt to give a false impression of accuracy. This is particularly true be- most of the specimens were surface finds. For these reasons, the specimens are related to igneous flow rocks or marker beds where possible, and then grouped into local faunas. cause Jeff Conglomerate DeFord (1958) assigned the basal conglomerate of the Tertiary section in the Vieja area to the Jeff Conglomerate of the Barilla Mountains, which are located on the east side of the Davis Mountains. The Jeff Conglomerate in the Vieja area was deposited prior to the volcanic activity. No fossils have been found in the Jeff Conglomerate either in the Barilla Mountains (Eifler 1951) or the Vieja area (DeFord 1958) or the Bofecillos Mountains (McKnight 1970), where the name has also been used. A variety of rocks of several different ages overlie the Cretaceous rocks in the Trans-Pecos area. For instance, there is a gradual transition from the late Cretaceous continental deposits to Paleocene continental deposits in the Tornillo Flat area in Big Bend National Park. Near Castolon in the southern part of the park the Alamo Creek Basalt (lateEocene, 42.2 m.y.) rests directly on the late Cretaceous Aguja Formation. In the Buck Hill and Agua Fria Quadrangles a “basal Tertiaryconglomerate" contains an early late Eocene (Uintan Wagonhound) vertebrate fauna and underlies a tuff dated 45.8 million years old. On Tornillo Flat where the stratigraphic section is fuller, the most likely correlative on the basis of lithology is the Big Yellow Member of the Canoe Formation (Maxwell et al. 1967). The Big Yellow, however, appears to be middle Eocene (Bridgerian) and therefore older. It cannot be demonstrated that the conglomerates at the base of the various Ter tiary sections are all part of one continuous body of rock so the name Jeff Conglomerate as applied beyond the Barilla Mountains 10 and the northern Davis Mountains should be used with caution. Gill Breccia The lowest volcanic rock of the ViejaGroup is the Gill Breccia (DeFord 1958). It has the maximum thickness of 300 feet (91 m) at its type section in Colmena Canyonbut thins to a few feet or pinches out completely in short distances. It varies in composition but in the vicinity of the mouth of Capote Creek it is a "brecciated to massive, light olive green to dark greenish-gray fine grained rock. Intimately associated with the breccia are blocks of Comanchean limestone, some as large as a three-story building." (Twiss 1972, p. 145). Attempts to date the Gill by potassium-ar gon resulted in two unsatisfactory dates of 75.0 ± 3.0 and 56.5 ± 3.8 and a much more ± geologically reasonable date of 40.0 2. The evidence to support the choice of the latter date is that it closely conforms to the date for the initiation of volcanic activityelsewhere in the west Texas volcanic field. Examples are the northern Davis Mountains and the Barilla Mountains (Parker and McDowell in press), the Agua Fria area, and Big Bend National Park. Colmena Formation The Colmena Tuff is the lowest formation in the Vieja Group that contains vertebrate fossils (fig. 4, secs. A, E, and fig. 6). Walton (1972), in the most recent description of the formation, divided it into three facies. The one that outcrops in the northern and central part of the area is an unfossiliferous sheet-like body dominated by Cretaceous calcareous rock fragments (CRF). This facies may be, in part, equivalent to the Jeff Conglomerate. To the south is a second facies characterized by volcanic rock fragments(VRF), and it is from this body of rock that the majority of vertebrate fossils have come, therefore, only this VRF facies will be treated at any length. In the southern area the VRF facies is overlain by a sparsely fossiliferous third facies composed mainly of volcanic glass shards and pumice fragments. Most of the Candelaria l.f. was found along and on both sides of the mouth of Capote Creek in the southern area. A few specimens have been collected from the Colmena in the vicinity of the type section (DeFord 1958) near Nixon Falls and one specimen about one mile northwest of the Dow House. No individual marker beds could be found within the Colmena. However, the petrologic studies of Walton (1972) have furnished valuable information on the facies within the Colmena and can be used to subdivide the approximately 250-foot (76 m) section at the mouth of Capote Creek. The Colmena, Chambers, and Capote Mountain were designated tuffs by DeFord (1958). Walton (1972) refers to them as for mations because the bulk of each consists of water-laid sediment. I agree with Walton, but the task of changing the names on diagrams and tables was too expensive. The following is a summary of Walton's de scription of the sedimentary petrology of the southern Colmena Tuff facies: "Sorted sandstones are common at the base of the Colmena. An excellent example of this occurs due east of the mouth of Colmena Canyon, on the ridge that forms the divide with Panther Canyon. The base of the Colmena is a sorted sandstone and pebbly sandstone unit some 20 feet thick, the middle of the formation is a sequence of massive, unsorted conglomerates about 200 feet thick and the top is a fine-grained unit rich in glassshards, about 25 feet thick. A similar situation obtains around the mouth of CapoteCreek." (Walton 1972, p. 54). The sorted sandstone beds at the base of the Colmena Tuff at the mouth of Capote Creek (fig. 6) are a dull red brown and have a banded appearance. They show faint cross- bedding and lamination, and Walton (1972) suggests they represent braided stream de posits. The middle unit of the VRF facies of the Colmena is interpreted as being a succession 11 Fig. 6.—Diagramaticsectionof ColmenaTuf at the mouth of Capote Crek's north side. LocationA is shownin figure 4. Faciesunits after Walton (1972). Distributionof vertebratefosils given in table 2. of lahars. An individual unit has a vertical size gradation from boulders at the base to silt and clay at the top. There is very little inter nal stratification. The middle 200 feet (61 m) of the Colmena consists of these kinds of sedi ments. Fossils are rare in this part of the sec tion; only a single tooth of an amynodont has been discovered and it was within the lower 20 feet (7 m) of the lahar facies. Walton calls the upper facies of the Col mena the glass facies (fig. 6) because 65 to 98 percent of the framework grains are shards and pumice. The informal unit, "Clabaugh's bonebed," TMM 40630, occurs in the upper or glass facies. Volcanic activity began in the Vieja area with the eruption of the Gill Breccia. A whole rock date of 40.0 ± 2 was obtained froma volcanic rock from the breccia. The sorted sandstones in the lower sedimentary unit of the Colmena Tuff contain a late Eocene (UintanMyton) fauna. The date of 40.0 m.y. for a rock underlying a Myton fauna is perhaps a little too young when compared with the date ± 41.2 1.4 m.y. for locality 20 at Badwater, Wyoming (Black 1974) that also contains a Uintan fauna, A date of 39.3 ± 0.8 for a tuff at the contact of the Halfway and LapointMembers of the Duchesne River Formation (McDowell et al. 1973) would also make the 40.0 date for the Gill seem young. In the appraisal of the age of the Colmena, as based on the rodents by Wood (1974), the Candelaria l.f. is "post-Myton in age but probably not very much later." CandelariaLocalFauna Apreliminary faunal list was given by Black and Dawson (1966) under the name Colmena fauna. Taxa that currently comprise the Candelaria l.f. and their stratigraphic positions are given in tables 1 and 2. No new elements have been added to the Candelaria l.f. that are useful for dating since the papers by Forsten (1971a), Wilson (1971a, 1974), and Wood (1974). The reports by Gustafson on the carnivores and Schiebout on the rhinos and tapiroids will be forthcoming. Of the material available for age evaluation, Epihippus cf. gracilis, according to Forsten (1971a), represents a southern popula- Table 1. Candelaria l.f.*, late Uintan. Dates refer to publica tions and are listed in the references. Taxon Reference Helix leidyi Pampe 1974 Oreohelix grangeri Pampe 1974 Alligator or crocodile Not identified Herpetological material Not identified PPeratherium Omomyid gen. et sp. indet Manitsha johanniculi Wood 1974 cf. eutypomyid gen. etsp. indet. Wood 1974 Ischryrotomus cf. / petersoni Wood 1974 Leptotomus leptodus Wood 1974 Prolapsus sp. Wood, pers. com. 8/75 Harpagolestes sp. Gustafson, in progress Epihippus cf. E. gracilis Forsten 1971a Sthenodectes australis Wilson 1977 Dilophodon n. sp. Schiebout, in progress Colodon cf. hancocki Schiebout, in progress Amynodon sp. Protoreodon petersoni Wilson 1971a Protoreodon pumilus Wilson 1971a Leptoreodonmarshi Wilson 1974 Toromeryx marginatus Wilson 1974 *ln addition a large amynodont lower jaw was loaned for studybut has, sincebeen lost.Itwastentativelyidentifiedby me as Megaiamydon and was last seen at the American Mu seum of Natural History, New York. Any assistance in its relocation will be greatly appreciated. Table 2. Distribution of Candelaria l.f. in Colmena Tuff Lower sorted facies Middle lahar facies (lower 50 ft.) (15m) Amynodonsp. Helix leidyi Oreohelix grangeri Upper Glass facies Alligator or crocodile (upper 25 ft.) PPeratherium Onomyid gen. etsp. indet Manitsha sp. Manitsha johanniculi Leptotomus leptodusIschryotomus cf. /. petersoni Prolapsus sp. Leptotomus leptodus Protoreodon petersoniProlapsus sp. Protoreodon pumilus Harpagolestes sp. Harpagolestes sp. Epihippus cf. E. gracilis Sthenodectes australis Dilophodon n. sp. Colodon cf. C. hancocki amnyodont, not identified Protoreodon petersoni Protoreodon pumilus Leptoreodon marshi Toromeryx marginatus 13 tion of E. gracilis. Protoreodon petersoni, a small protoreodont, is found in the Candelaria l.f. but not in the overlying Porvenir I.f. The very small P. minimus, however, occurs at McCarty's Mountain in the Chadronian of Montana, so it is possible that a comparablysmall form will be found in the Porvenir l.f. The type of P. petersoni is from the Uinta C, Myton Pocket, Uinta County, Utah. In Texas, Protoreodon pumilus, a larger protoreodont, extends from the Candelaria l.f. to the Chadronian Porvenir l.f. in the lower part of the Chambers Tuff. The latter occurrence is the highest, to my knowledge, and the only one where Protoreodon pumi/us is associated with a Chadronian fauna. In Utah, P. pumilus is found as low as Uinta B. Emry (1975) reviewed the occurrences of Protoreodon at Beaver Divide and assigns them to the Uintan. Toromeryx is endemic and therefore of no assistance for correlation. According to Wood (1974, p. 104): "Four of the six rodent specimens from the Colmena Tuff (Candelaria local fauna) are referred to species previously known from the late Eocene Uinta Formation of Utah (Leptotomus leptodus and Ischyrotomuspetersoni, each occurring in both the Wagon- hound and Myton local faunas). The other two specimens are useless in correlation. Leptotomus leptodus is more advanced than the material of the same species from the Myton. The rodent evidence, then, suggests that the Colmena (Candelaria 1.f.) ispost-My ton in age but probably not very much later." Buckshot Ignimbrite The Buckshot Ignimbrite (figs. 5,7, 8) is a very important marker throughout the Vieja area. It is easily identifiable from hand specimens, and its upper surface in many areas is covered with hemispherical mounds properly called tumuli but informally known as "blister cones." These make the Buckshot easy to identify at a distance. The ignimbrite is more resistant to erosion than the Colmena below or the Chambers above and therefore caps mesas and buttes, and forms prominent dip slopes. Anderson (1975), in a recently completedstudy of the Buckshot, states (p. 13): "The Buckshot Ignimbrite is a zoned ash- flow sheet composed of one simple coolingunit and overlain by an air-fall ash depositthat is believed to be a product of the same explosive eruption. The Buckshot is exposedin a narrow belt extending 75 km. from north to south and 15 km. from east to west . . Its color varies from grayish brown to . dark. reddish brown, the reddish brown most often found on weathered surfaces In outcrop . ... or hand specimen, easily recognizable constituents are colorless phenocrysts of alkali feldspar, glass shards, fragments of intermeditate volcanic rocks and various secondaryminerals filling cavities in the rock." The Buckshot provides a time plane because as a single cooling unit it must have been deposited very rapidly. For this reason a postassium-argon date was felt to be very important. Three dates are listed by Wilson et al. (1968) for the Buckshot Ignimbrite : ±± 34.7 2.0, 35.2 2.3 and 38.6 ± 1.2. Of these, the 38.6 was chosen as the best date because it is older than the 36 m.y. dates on the Bracks Rhyolite which is stratigraphicallyhigher. This choice has been substantiated bythe whole suite of dates on flow rocks in the northeastern Davis Mountains, which are 36 37.5 m.y. (fig. 3, sec. 7, table 15) and - which, like the Bracks Rhyolite, overlie Oligocene vertebrates (Sellards et al. 1933, p. 805). The date of 38 m.y. for the beginning of the Chadronian (Berggren and van Couvering 1974) now seems well established. In the north-central and western parts of the Vieja area the Buckshot pinches out and the Chambers Tuff concordantly overlies the Colmena. The two are sufficiently distinct lithologically to be easily distinguished without the Buckshot. Chambers Tuff The Chambers Tuff was defined by DeFord 14 feet300 lower the of interval stratigraphic The FMNH. the at labels top to Ignimbrite Buckshot from section 7.—Diagrammatic Fig. Die- Arroyo to northMesa Carr from extending Chambers the of those areunits informal 4.The figure in given Location Cliff. Bigof a is 40492 TMM l.f. Porvenir 5. the table of most mount," of “red source as the is known chiocholocality specimen onusedareand notes field his in Patterson Bryan by given Chambers the of part lower the of section B.—Diagrammatic Fig. position showing area Vieja southern the in Springs Adobe near Tuff l.f. Porvenir the of elements containing unit fossiliferous of (1958) as "the strata between the top of the Buckshot Ignimbrite and the base of the Bracks Rhyolite." The term tuff is misleading because most of the Chambers consists of water-laid sediments. Individual beds rangefrom coarse conglomerate lenses containingbone fragments, to units of sand, silt, or clay. The conglomerates contain cobbles of fossiliferous Cretaceous limestone and fragmentsof igneous rock. The conglomerates are usually gray but the finer grained units are more colorful and can be brick red, salmon red, pink, greenish brown or dark brown. The lower part of the formation is well exposed at Big Cliff and the surrounding area (figs. 4,7). Walton (1972, 1975) has studied the sediments of the Vieja Group in detail and has made the very valuable contribution of recognizing and tracing lower and upper marker beds within the Chambers (figs. 7,8). Although he does not map the marker beds, he very carefully describes their color, lithology, stratigraphic position, and distribution: "The lower of these marker beds is a tuff bed, 6 to 30 feet thick that crops out 1GO- 175 feet above the Buckshot... it is the same bed described as 'white volcanic dust' at the Big Cliff mammal locality by Stovall (1948, see also Mankin, 1955). The bed is white to pale orange or pale green and usually is less resistant to weathering than beds above and below it. ." (Walton 1972, p. 79). . The upper marker bed of Walton (1972, 1975) had been previously recognized by students mapping in the area. It was informally named the Chambers red siltstone lentil by Peterson (1955) but the name was changed when DeFord formally described the Chambers Tuff. Twiss (ms. map of Vieja area) maps this unit as the "Rooney siltstone lentil." It is a hard, resistant, brick red unit that forms dip slopes and caps buttes and mesas where the Bracks has been removed. Walton (1972, p. 83) describes it as follows: "Eighty percent of it is silt-to-fine sand- sized, well sorted, euhedral, overgrown sanidine. Grains of volcanic quartz, slightly coars er than the sanidine, total about 5% of the rock. It is cemented by fine-grained quartzand calcite that amount to about 15 to 20 percent of the rock. Evidence that the upper marker bed itself is a tuff is that it uniformly covers such a large area; is of quitedifferent composition than any other unit of the Chambers section; and, though well sorted is marked by no sedimentary structures, unlike most well sorted sedimentary rocks." The "upper marker bed" of Walton (1972,1975) or the "Rooney siltstone lentil" of Twiss (ms. map) holds up Dunagan Mesa and Carr Mesa that lie to the east and south of Big Cliff. Mankin's thesis (1955) contains stratigraph ic sections and correlations of the lower part of the Chambers from the field notes of Bryan Patterson that were made in 1946. Patterson refers to what is now the Buckshot as the "basal Vieja lava" and Walton's lower marker bed as "Yellow white tuff, the white layer." Patterson uses the "white layer." (fig. 7) to correlate from Big Cliff north to Soldier Camp Flat. The stratigraphic interval between the Buckshot and the lower marker bed is Patterson's "Big Red Horizon" and an interval up to approximately 100 feet above the lower marker bed he calls the "Blue Cliff Horizon." Walton (1972, p. 79) believes the lower marker bed is an ash flow tuff which he recognizes from the southern to the northern extremes of the Vieja area, a distance of about 40 miles. The lower marker bed is especiallyuseful in the Big Cliff-Soldier Camp Flat area because most of the Porvenir l.f. came from either the hundred feet of strata just below or above it. The lower marker bed has not been submitted for potassium-argon dating, but it would be a most useful one and, if the sample were suitable, would form an excellent time plane just above the Buckshot. However, it might well be that the dating method would not be able to resolve the time difference between it and the Buckshot. The time span must have been short because I am unable to detect evolutionary changes within the Porvenir l.f. which extends from just above the 17 Buckshot to about 100 feet above the lower marker bed. Figure 7 is a measured section of the lower part of the Chambers Tuff from the Buckshot Ignimbrite to the top of Big Cliff (fig. 4, sec. B) and includes the lower marker bed. The greatest variety of taxa of the Porvenir l.f. were found in the lower part of the Chambers Tuff. All but one of the TMM localities from this part of the section were isolated surface finds. The single exception is TMM 40492, "red mound," where a concentration of material listed in table 5 was found. Figure 8 is a measured section of the lower part of the Chambers Tuff from the Buckshot to the lower marker bed near Adobe Spring south of Capote Creek with the stratigraphic positionof additional elements of the Porvenir l.f. Adobe Spring (fig. 4, sec. C) is labeled "Mexican Spring" on the map in Mankin (1955). The upper boundary of the Chambers Tuff is placed at the base of the Bracks Rhyolite(DeFord 1958).. The Bracks, unfortunately, is not present in the area north of Soldier Flat. Here, uppermost Chambers and the lower Capote Mountain are so similar lithologicallythat in places it is difficult to distinguishthem. A third or uppermost marker bed, however, was used by Schulenberg (1958). He described it as follows: "The Chambers-CapoteMountain contact or the horizon of the Bracks, has been approximated in the Porvenir area by mapping the upper contact of a moderate red sandstone which immediatelyunderlies the Bracks where the Bracks is present" (Schulenberg 1958, p. 49). Two important localities, Reeves bone- bed and Chalk Gap Draw, occur in the area where the Bracks is absent. The red sandstone is present in the Reeves bonebed area and is the uppermost sandstone in figure 9. A measured section was not made in the Chalk GapDraw area but the specimens from there and Reeves bonebed form the Little Egypt l.f. and are at approximately the same level. Porvenir Local FaunaTable 3 includes all taxa of the Porvenir l.f. with the originaland some important references to those taxa. Tables 4-7 list parts of the Pon/enir l.f. from specific stratigraphic levels or geographic areas. Table 4 lists only those taxa of the Porvenir l.f. that were found in the lower 100 feet of the Chambers Tuff or beneath the lower marker bed. This unit is called the "BigRed horizon" in the field notes of Bryan Pat terson at the Field Museum of Natural History, Chicago. It is also the "main fossil level" of Stovall (1948). Taxa from the TMM localities "at the foot of Big Cliff," "north of Big Cliff," and 40263 for the skull of Hemipsalodon are also included in table 4. Table 5 lists those taxa of the Porvenir l.f. that were found at the locality known as "red mound north of Big Cliff." The rocks at this locality are less than five feet below the lower marker bed. Table 6 lists taxa included in the Porvenir l.f. that were found between 0 and 88 feet above the lower marker bed. This unit is the "Blue Cliff horizon" in the field notes of Bryan Patterson at the FMNH. Table 7 lists only those taxa of the Porvenir l.f. that were found near Adobe Springs in the southern part of the Vieja area. Little Egypt Local Fauna Table 8 lists all the taxa and original references to the Little Egypt l.f. Table 9 is restricted to taxa from Reeves Bonebed, and Table 10 to taxa from Chalk Gap Draw. Capote Mountain Tuff In the northern part of the Vieja area the Capote Mountain is a fine-grained, vitric, tuffaceous sandstone and siltstone. Unfortunate ly, there are no marker beds in the CapoteMountain along Walker Creek. The bulk of the Capote Mountain in this area is a succession of pinkish gray to dark brown spheroidally weathered, fining upward siltstone and sandstones. Very faint cross-bedding is found in some siltstones but distinct sedimentary structures are rare. Clay pebbles form lenses of conglomerate and are irregularly distributed in the sandstone and siltstone. The stratigraphic position of the Airstrip l.f. is estimated as 570 feet above the Bracks Rhyolite. Although most of the specimensfrom the Airstrip l.f. were found within a 18 Chambers the of part upper of section 9.-Diagrammatic Fig. marker sandstone salmon-red the Bonebedto Reeves from Tuff Rhyolite. Bracks the of basethe marks thatbed Table 3. Porvenir local fauna, early Chadronian [Table 3, continued] Taxon Planorbis sp. Mesodon ("Helix") leidyi Lacertilia (not seen) Herpetologicalmaterial, unstudied Rooneyia viejaensis Geolabis n. sp. Leptictis douglassi Apternoduscf. A. brevirostris L ep totomus gigans Microparamys perfossus Ischryotomus cf. /. petersoni Manitsha joahniculi Ischryomys blacki Pseudocylindrodon neglectus Pseudocylindrodon texanus Ardynomys occidentalis Adjidaumo cf. A. minutus Viejadjidaumo magniscopulicf. Viejadjidaumo, sp. indet. Aulolithom ysbouni tes Yoderimyslustrorum Eutypomys inexpectatuscf. Eutypomyid, gen. et sp. indet. cf. Simimys, sp. indet. IschnognathussavageiHemipsalodon n. sp. Hyaenodon sp. Miacis n. sp. Daphoenussp. Haplohippus texanus (Mesohippus cf. M. bairdi) Mesohippustexanus Mesohippussp. Menodus baked Teleodus sp. Colodon n. sp. Caenopus sp. Hyracodon sp. Metamynodon sp. Schizotheroides jackwilsoni Brachyhyops wyominensis Reference Stovall 1948 Goldich & Elms 1949, L. S. Russell; Pampe 1974; Stovall 1948 Wilson 1966; Hoffer & Wilson 1967; Szalay 1976; Jerison 1973; Szalay & Wilson 1976 Lillegraven, pers. com. 1972 Novacek 1976 Novacek 1976 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Wood 1974 Stovall 1948 Gustafson, in progressGustafson, in progressGustafson, in progressGustafson, in progressMcGrew 1953 Stovall 1948; but see McGrew 1971; Forsten 1971a, b; Clark et al. 1967 McGrew 1971; Forsten 1971a,b McGrew 1971; Forsten 1971a, b Stovall 1948; Wilson 1977 fide Clark et al. 1967; Wilson 1977 Stovall 1948=Co/oc/or7 n. sp., Schiebout, in progressSchiebout, in progressStovall 1948; Schiebout, in progressSchiebout, in progress Schiebout 1977 Wilson 1971b Taxon Archaeotherium cf. A mortoni Protoreodon pumilusAgriochoerusantiquus Aclistomycter middletoni Mercoidodontidae gen. et sp. indet. Heteromeryx disparPoabromyluskayiPoabromylus minor Oromeryx sp. Eotylopus cf. E. reedi Hidrosotherium transpecosensisLeptomeryx defordi Reference Wilson 1971b Wilson 1971a identified as Merycoidodon cuibertsoni, M. gracilis, and Mesohippus texanus in Stovall 1948. Wilson 1971a Wilson 1971a Wilson 1974 Wilson 1974 Wilson 1974 Wilson 1974 Wilson 1974 Wilson 1974 Wilson 1974 Table 4. Taxa found in the lower 100 feet of Chambers Tuff or beneath the lower marker bed; Porvenir l.f. Herpetological material, unstudied Leptotomusgigans Ischyrotomus cf. petersoni Eutypomys inexpectatus Pseudocylindrodon neglectus Manitshajohanniculi Hyaenodon sp. Hemipsalodon n. sp. Ischnognathussavagei Haplohippus texanus Mesohippus texanus Mesohippussp. Menodus bakeri Teleodus sp. Colodon n. sp. Hyracodon sp. Metamynodonsp. Coenopus sp. Schizotheroides jackwilsoni Brachyhyops wyomingensis Protoreodon pumilus Agriochoerus antiquus Heteromeryx dispar Poabromylus minor Poabromylus kayi Leptomeryx defordi 20 Table 5. The following taxa included in the Porvenir l.f. Table 7. The following taxa included in the Porvenir l.f. were found just below (less than 5 feet) "the lower marker were found at localities in the southern part of the Vieja bed" of Walton (1972) in the Chambers Tuff at locality area near Adobe Springs, 25-42 ft. above the Buckshot "red mound, north of Big Cliff," TMM 40492 Ignimbrite. Gastropods, unstudied Herpetological material, unstudied Geolabis n. sp. Apternoduscf. brevirostris Aulolithomysbounites Viejadjidaumo magniscopuliMicroparamys perfosuscf. Viejadjidaumo sp. indet. Adjidaumocf. minutus Yoderimys lustrorum cf. Leptotomus gigans PseudocylindrodonneglectusMesohippustexanus Colodon n. sp. Hyracodon sp. Agriochoerusantiquus Leptomeryx defordi Table 6. The following taxa included in the Porvenir l.f. were found between 0-88 feet above the "lower marker bed” of Walton (1972) in the Chambers Tuff. This unit is the "Blue Cliff horizon" in the field notes of Bryan Patterson at the Field Museum of Natural History, Chicago. Gastropods,unstudied Herpetological material, unstudied Rooneyia viejansisLeptictis douglassiHyaenodon sp. Mesohippustexanus Teleodus sp. Colodon n. sp. Hyracodon sp. Metamynodon sp. Caenopus sp. Protoreodon pumilusArchaeotherium cf. mortoni Agriochoerus antiquus Oromeryx sp. Eotylopus cf. reedi Hidrosotherium transpecosensis Leptomeryx defordi five-foot interval, occasional turtle carapaces and isolated bones were found lower and higher in the section. The bones are usually found in nodules that seem to be the result of the spheroidal weathering. Walton (1972, p. 85) says that: Gastropods, unstudied Herpetological material, unstudied Leptotomus gigansMicroparamys perfossusIschryotomusblacki Aclistomyctermiddletoni Table 8. Little Egypt local fauna, Chadronian Reference Taxon Helix leidyi Pampe 1974 Leptictis wilsoni IMovacek 1976 Manitshajoahanniculi Wood 1974 Cylindrodon fontis Wood 1974 Pseudocylindrodon neglectus Wood 1974 Pseudocylindrodon texanus Wood 1974 Ardynomys occidentalis Wood 1974 Aulolithomys cf. A. bounites Wood 1974 Eutpomys inexpectatus Wood 1974 Hyaenodon sp. Gustafson, in progress Miacis n. sp. Gustafson, in progress Daphaenocyon sp. Gustafson, in progress Fetid gen. et sp. indet. Gustafson, in progress Mesohippustexanus Menodusbaked Wilson 1977 Colodon n. sp. Schiebout, in progress Hyracodon sp. Schiebout, in progress Schizotheroidesjackwilsoni Schiebout 1977 Archaeothenum cf. A. mortoni Wilson 1971b Agriochoerusantiquus Wilson 1971a Bathygenys reevesi Wilson 1971a Merycoidodon dunagani Eotylopus cf. E. reedi Wilson 1974 “Spheroidal weathering is common in poorly bedded intervals rich in diagenetic clayand clinoptilolite shard pseudomorphs in which the clay expands as it is wetted byrainfall and contracts when dried." Nodules that contain skulls often had the main line of fracture passing longitudinallythrough the tooth row. On some specimens a concentration of very hard hematite next to the bone or dentine made cleaning very difficult. The tuffaceous sandstones are consolidated and not suitable for washing. In an at 21 tempt to increase the collection 200 nodules were broken open but not a single one contained bone. In addition, an area of about 400 square feet of nodules was raked over. The idea was that at least as many nodules should occur with the bone exposed on the lower side. Only one additional specimen was found within this area. Table 11 lists the taxa in the Airstrip l.f. Table 9. The following taxa included in the Little Egypt l.f. were collected from or very close to "Reeves Bonebed," in the upper part of the Chambers Tuff. Gastropods, unstudied Herpetological material, unstudied Manitsha johanniculiLeptictis wilsoni Cylindrodon fontis Pseudocylindrodon neglectusArdynomys occidentalis Aulolithomys bounites Eutypomys inexpectatusHyaenodon sp. Miacis n. sp. Daphyaenocyon sp. Felid, gen. et sp. indet. Mesohippus texanus Menodusbaked Colodon n. sp. Hyracodon sp. AgriochoerusantiquusBathygenysreevesi Merycoidodon dunaganiEotylopus cf. E. reedi Table 10. The following taxa included in the Little Egypt l.f. were collected from localities in Chalk Gap Draw in the upper part of the Chambers Tuff. Gastropods, unstudied Herpetological material, unstudied Pseudocylindrodon texanus Menodus baked Schizotheroides jackwi!soni Archaeothedum cf. mortoni Merycoidodon dunagani Eotylopus cf. reedi Vieja Group Undifferentiated The name Vieja Group undifferentiated is used where a volcanic unit and one or more of the marker beds are absent either by pinch- out or faulting, and it is not possible to determine the boundary of a sedimentary unit. This must be done only at the Ash Spring locality. Although the rodents suggest that the Ash Spring l.f. is the youngest, its exact stratigraphic relationship to the beds that contain the other local faunas cannot be determined because of faulting. The Ash Spring locality was studied and collected in 1966 and 1967 by John M. Harris. It was discovered and mapped by Bridgesand Dasch and first described by Bridges(1958). Bridges placed the sediments at the Ash Spring locality in the upper part of the Capote Mountain but this assignment cannot be proven because of the absence of the marker beds or marker flows in the area. The Ash Spring locality is the northernmost and westernmost one in the Vieja area. The sediments are poorly consolidated and their color, a light pinkish gray, resembles the Miocene and younger bolson fill. The location (N3o°37'W 104°53'30") is shown on figures 1 and 4H, and on Bennett Ranch Quadrangle, Texas, USGS 7.5' series (topographic). Bridges (1958) and Harris (1967a) give measured sections of the Ash Spring area. Table 11. The following taxa were found at or very close to TMM 40504 approximately 570 ft. above the Bracks Rhyolite in the CapoteMt. Tuff, Airstrip 1.f., Chadronian. Taxa References Herpetological material, unstudied Subsumus candelariae Wood 1974 Pseudocylindrodon cf. neglectusJaywilsonomys aff. pintoensisArdynomys occidentalis Hyaenodon sp. Hesperocyon sp. Hyracodon sp. Bathygenysreevesi Limnenetes cf. platyceps?Prodesmatochoerus cf. meekae Wood 1974 Wood 1974 Wood 1974 Gustafson, in progress Gustafson, in progress Schiebout, in progress Wilson 1971a Wilson 1971a Wilson 1971a Poebrotherium franki Wilson 1974 Poebrotherium sp. Hypisodus cf. minimus Wilson 1974 Wilson 1974 22 Ash Spring Local Fauna —Harris (1967b) lists the taxa that were known from Ash Spring as of that date. A revised list is givenin table 12. Garren Group The upper member of the Hogeye Tuff of the Garren Group has produced a small collection of vertebrate fossils. The locality was found by Underwood and, although it is not labeled on his geologic quadrangle map (Underwood 1963) it is in the upper member of the Hogeye Tuff (Underwood 1963, p. 18) and is located near his measured section 4. The Garren Group interfingers on the north with the Vieja Group and the Hogeye Tuff is in part equivalent to the Chambers Tuff (fig. 3). At the time Underwood published his map and text I stated that the faunule from the Hogeye correlated with the Ash Spring l.f. I now believe it much more probablethat the faunule from the Hogeye Tuff correlates with the Porvenir l.f. Taxa identified from the Hogeye Tuff are as follows; Titanothere gen. et sp. indet., Mesohippus texanus (in McGrew 1971), Agriochoerus antiquus, and Leptomeryx sp. Table 12. Ash Spring local fauna, Chadronian Taxon Reference Gastropids,unstudied Herpetological material, unstudied Avian material, unstudied Titanotheriomys veterior Harris 1967a, b; Wood 1974 Pseudocylindrodon cf. P. texanus Wood 1974 Meliakrouniomys wilsoni Harris & Wood 1969; Wood 1974 Felid, gen. et sp. indet. Harris 1967a Rhinoceratid, gen. et sp. indet. Harris 1967a Toxotherium cf. T. woodi Harris 1967a Archaeotherium cf. A. mortoni Wilson 1971b Merycoidodontid, gen. et sp. indet. Harris 1971a; Wilson 1971a Poebrotherium franki Wilson 1974 Oromerycinae,gen. et sp. indet. Wilson 1974 Hypisodus cf. H. minimus Wilson 1974 Prietos Formation A small area of volcanic flows and intercalated sediments occurs 35 miles northwest of Ojinaga, Chihuahua, Mexico. The area was mapped by Grant H. Heiken and Ismael Ferrusquia- V.in 1965. Ferrusquia-V.(1969) described the Rancho Gaitan l.f. and published a geologic map and section of the area. Heiken (1971) described the petrology, defined the Prietos Formation, and tentatively identified the Brite or Mitchell Mesa Ignimbrite. Although the Rancho Gaitan area is not partof the Vieja area proper, the local fauna and the stratigraphic section are more closely related to the Vieja than to the folded marine Cretaceous that bounds the area to the north, west, and south. Rancho Gaitan Local Fauna —This local fauna is a very important one in that it contains some taxa that have not been discovered the Oligocene on the east side of the Rio Grande, for example, Jaywilsonomys, Protoreodon petersoni, Agriochoerus maximus, and Hypertragulus heikeni. It furnishes additional valuable information for correlation of the southern Chadronian faunas to those of both Montana and Wyoming and also to those of Southern California. Table 13 lists the taxa assigned to the Rancho Gaitan l.f. Chronostratigraphy, Vieja Group Radiochronology One of the long-term purposes of the project in west Texas is to tie the stratigraphic position of vertebrate fossil faunas to potassium-argon dated rocks. Samples were collected and submitted to various laboratories when funds were available. The first results were incorporated in Evernden et al. (1964). Additional dates were published in Wilson et al. (1968) for the Vieja area; and in Maxwell et al. (1967) and Maxwell and Dietrich (1970, 1972) for Big Bend National Park. Potassium-argon dates for the Buck Hill Group are found in Gilleland et al. (1969) and in Maxwell and Dietrich (1972). 23 Table 13. Rancho Gaitan local fauna, Early Chadronian Taxon Reference Gastropods, unstudied Herpetological material, unstudied Mytonomys gaitania Ferrusquia-V. & Wood 1969 Jaywilsonomysojinagaensis Ferrusquia-V & Wood 1969 Jaywilson omys pintoensis Ferrusquia-V & Wood 1969 ?Brontops cf. B. brachycephalusPHyracodon sp. Rhinoceratoidea, gen. et sp. indet. Protoreodon petersoni Agriochoerusmaximus Bathygenysa!fa* Merycoidodontidae, gen. et sp. indet. Hypertragulus heikeni cf. Leptomeryx sp. Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 Ferrusquia-V 1969 ‘Probably B. reevesi More recently, several dates have been run byParker (1976) and Parker and McDowell (1973, and in press) for igneous rocks in the northern Davis and Barilla Mountains (fig. 3, table 15). The igneous rocks dated by Parker and McDowell overlie the Huelster Formation from which came the Hyracodon tooth, TMM 41256-4, collected by C. L. Baker, identified by R. L. Stirton, and mentioned first in Sellards (1933). Baker also collected a flora from the Huelster Formation that was described by Berry (1919) and recently reviewed by Axelrod and Bailey (1975). Hyracodon is usually considered an Oligoceneform and is found in the Porvenir l.f. in the Vieja area. This adds support to the proposal in Wilson et al. (1968) that a date of 38.6 for the age of the Buckshot Ignimbrite is the most logical of the ones presented for the base of the Oligocene. The oldest igneous flow rocks of the Tertiary are in the Canoe Formation (Bridgerian) in Big Bend National Park but in spite of several attempts no samples suitable for datinghave been collected. The earliest date in the Big Bend area (47 m.y.) is on a tuff that directly underlies a very early Uintan Whistler Squat l.f. of the Agua Fria area that is in turn overlain, but not directly, by a biotite rich tuff dated at 42.9 m.y. These dates correlate closely with 41.2 ± 1.4 given by Black (1969) for tuffs associated with vertebrate faunas of Uintan age in the Badwater area, Wyoming. The Texas dates correlate closely with those given by Mauger (undated report) for Utah and Wyoming and are certainly additional evidence to support the proposed recalibration of the Eocene by McKenna et al. (1973). More data on these dates will be given in Part II of this report. The dates recently run by Parker and McDowell (1973 and in press) on the volcanic rocks associated with vents in the Paisano Pass area and Chinati Peak add support to the growing mass of evidence that volcanic activity in the Big Bend area began about 47 m.y. ago, reached its peak from 38-26 m.y. and gradually died out from 22-17 m.y. ago. Biochronology Candelaria Local Fauna-Table 14 shows the stratigraphic distribution of the currentlyidentified taxa from the five local faunas in the Vieja area. The oldest or Candelaria l.f. is not large in terms of specimens and taxa but is composed of a new association of taxa. Prolapsus is now known in the Candalaria l.f. whereas it was first discovered in the AguaFria area (Part 11, this study) and was identified by Wood (1973). In the Agua Fria area it is associated with a rodent fauna that Wood (1973) dated as Bridgerian, so it is now known to extend from the Whistler Squat l.f. to the Candelaria l.f. Manitsha extends from the Candelaria through the Porvenir to the Little Egypt local faunas. It was previouslyknown only from the Brule Formation of the Slim Buttes area of South Dakota; however, the locality and age of the type is uncertain (Lillegraven 1970). Leptotomus leptodus, late Eocene in Utah, is present in the Candelaria l.f. and with Harpago/estesEpihippus, Sthen, odectes; Protoreodon petersoni, and Leptore odon marshi certainly form a late Eocene as sociation. The work in progress by Schiebout on Dilophodon and Colodon cf. hancocki do not contradict this conclusion. The 40 m.y. date from the stratigraphically 24 lower Gill Breccia means that the Candelaria l.f. is approximately that old (Wilson et al. 1968). The date of 39.3 by McDowell et al. (1973) for an ashy siltstone at the contact of the Halfway and Lapoint members of the Duchesne River Formation would place the Halfway l.f. in the late Eocene. Epihippus intermedius is the only taxon in the Halfway 1. f. known elsewhere, and according to Forsten (1971a), "The measurements on P4-M2 from the Candelaria l.f. fall among the observations on the hypodigm of E. gracilis, and in size to a group consisting of material of that species and of E. intermedius." This correlation partly supports the interpretation givenby McKenna et al. (1973) in which the Brennan Basin (Randlett) and Dry Gulch Creek (Halfway) Member and the Candelaria l.f. are placed in the Eocene. The most common form in the Candelaria l.f. is Protoreodon petersoni; followed by Epihippus. Less common are Protoreodon pumiius and Leptogragaius, with other taxa known only from single specimens. Last appearances in the Candelaria l.f. in the lower sorted facies include Harpagolestes, Epihippus, Di/ophodon, and Leptoreodon, whereas Prolapsus makes its last appearancein the upper glass facies. Toromeryx is an endemic form perhaps ancestral to Heteromeryx. Porvenir Local Fauna This unit contains the largest assemblage of taxa in the Vieja area and is the only one usually referred to in older literature as the "Vieja Fauna." The discovery of a succession of local faunas in stratigraphic superposition necessitated the assignment of local fauna names to collections made within limited stratigraphic intervals. Thus, the Porvenir l.f. includes forms found in the Chambers Tuff above the Buckshot Ignimbrite and below the upper marker bed. The assemblage has the association of Mesohippus with Protoreodon and Poabromylus, which, although now known to occur in other Chadronian faunas, was first recognized by Wilson (1971a, 1974).Perhaps more significant, however, is the association of Hemipsalodon, Brachyhyops, and a primitive species of Leptomeryx. Such an association occurs within the "Lower banded zone" of the Little Lone Tree Gulch sections at Flagstaff Rim, Wyoming (Emry, pers. com., and Emry 1973). The lower limit of sedimentary rocks of the Chambers Tuff and the limit of the possibilityof finding members of the Porvenir l.f. is at the stratigraphic contact with the top of the Buckshot Ignimbrite. The first appearance, therefore, of the members of the Porvenir l.f. is the result of the beginning of sedimentaryaccumulation following a local volcanic eruption and is not necessarily of regional significance. The total assemblage of the Porvenir l.f. enclosed by potassium-argon dates is regionally significant. In addiition, the appearance of new taxa higher in the section and the order of their appearance is significant although the proof of an appearance as "the first" is good only until an earlier one is found. The faunal lists in table 14 do not give the frequency of occurrence of the various taxa. Not all data are currently at hand but table 16 is a census of numbers of specimens of some of the medium-sized herbivores in the Porvenir l.f. The samples were taken independently at different times. The first was taken by Langston, McAnulty, and Savageunder Stovall in 1938 and 1940 when approximately one week was spent in the field; the second, by Patterson and Quinn for the Field Museum of Natural History when three months were spent in the field. I have collected from the same area for a total time of approximately two months accompaniedby various colleagues and field assistants. All of the material from the Porvenir l.f. was collected by surface prospecting because the highly consolidated matrix cannot be washed. All three collections show a similarity of frequency of genera (I do not have figureson the collection of Leptomeryx at the FMNH but know it to be larger than five). Mesohippus and Agriochoerus are by far the most common genera in collections and none of the specimens of these two genera were found in quarry situations. Leptomeryxis common but this is a much smaller form. 25 Heteromeryx, Protoreodon, and Poabromylus are not rare in the total collection. Merycoidodonts, on the other hand, are rare. Ac/istomycter is found at a similar stratigraphic position in the southern Vieja area. A comparison with Clark et al. (1967, fig. 25) for the genera of the Chadron Formation shows that Mesohippus is common in the Ahern and Crazy Johnson but becomes rare in the overlying Peanut Peak. Agriochoerus is rare in the Ahearn and Crazy Johnson and not known but probably existed during the time of deposition of the Peanut Peak. Unfortunately, Clark et al. (1967) did not study the small selenodont artiodactyls from the Chadron Formation of South Dakota. Through the courtesy of Dr. John Clark I was able to examine the small artiodactyls from the Chadron Formation in the FMNH. To describe that large and valuable collection is beyond the scope of this study but I can report on certain conclusions that are pertinent to this report. I hope that the Chicago material will be reported on fully in the not-too distant future. Leptomeryx is the most common small artiodactyl in the FMNH collection from the Chadron Formation; 44 specimens were borrowed and this was not all of the Leptomeryx specimens in that collection. In addition, 37 partial dentitions or isolated teeth of Poabromylus were examined. Eleven specimens of Poebrotherium, two specimensof Heteromeryx, and one of Eoty/opus were certainly identified. In addition and very important were three specimens labeled Pentacemylus and one labeled PCamelodon. A comparison of the specimens of Pentacemyluswith casts of dentitions from the Myton of Utah showed no difference. Pentacemylus is now known from the Myton, is listed by Andersen and Picard (1972) from the Lapoint, and is also present in the Chadron Formation. The specimens so identified are FMNH PM 22760, P4-M2; 22759, M3; 22758, 7P2-P3 The specimen number of the PCamelodon is FMNH PM 22754. More detailed work is needed on the small artiodactyls of the Chadron Formation of South Dakota and my only purpose here is to point out their presence in the northern fauna. Nonetheless, the data in tables 14 and 16 are useful for interpreting faunal succession and with the control of potassium-argon dates it is possible to draw preliminary conclusions. The date of 38.6 m.y. for the Buckshot Ignimbrite beneath the Porvenir l.f. and the large number of dates run by Parker and McDowell given in table 16 and the 36.5 date on the Bracks Rhyolite bracket the time spanof the Porvenir l.f. In this interval the most commonly found medium-sized herbivores in Texas are Mesohippus and Agriochoerus. Beneath the Buckshot Ignimbrite ii the Col / mena Tuff and beneath it the Gill Breccia. volcanic rock from the Gill Breccia was dated at 40 m.y. (Wilson et al. 1968). This brackets the Candelaria l.f. between 40 and 38 m.y. The most commonly found herbivores in the Candelaria l.f. are Protoreodon (13 specimens) and Epihippus (four specimens). This succession agrees in part with Clark et al. (1967, fig. 24). They list an association of Mesohippus and Agriochoerus overlying Epihippus, Diplobunops, and Poabromylus. They also include Te/eodus but the titanotheres from this part of the section are sorely in need of restudy. Clark et al. (1967) also include Diplobunops and Poabromylus as associates in the "index fauna" with Teleodus and Epihippus. Diplobunops has not yetbeen found in Texas, so its association cannot be verified. It may be that Diplobunops either did not inhabit the Texas area or that Diplobunops was present in an earlier association not found in the Vieja area. Poabromylus, on the other hand, is present in Texas and is now known from the Lapoint and the Chadronian, so is not a useful part of an "index fauna" to identify the early Duchesnean of Clark et al. (1967, fig. 24). McDowell et al. (1973) give a date of 36.3 ± 0.7 for an unnamed tuff three feet below the top of the Ahearn Member of the Chadron Formation. This date correlates as closely as can be expected with the dates of 36.8 and 36.5 ± 1.2 for the Bracks Rhyolitethat overlies the Chambers Tuff. Ash B of the Flagstaff Rim section lies above the occurrence of Brachyhyops and Hemipsalodonand was dated by Evernden et al. (1964) as 26 Table 14.—Stratigraphic occurrence of currently identified taxa in the local faunal succession of theVieja area. Figures representpotassium/argon dates that show the best fit to the superposition and composition of the faunas. 40.0 is from a boulder within the Gill Breccia stratigraphically beneath the Candelaria l.f. (Wilson et al. 1968); 38.6 is from the Buckshot Ignimbrite (Wilson etal. 1968);36.8isfromtheBracksRhyolite(Wilsonetal. 1968);31.4isthecurrentlyacceptedaveragedatefortheBrite Ignimbrite which is the equivalent of the Mitchel Mesa Rhyolite to the south (McDowell, personal communication 1977). Taxa of certain identification in the Rancho Gaitan l.f. are shown as G, although their position with respect to the date on the Bracks is based on correlation only. Position of the Ash Spring l.f. is based on evolutionary position of taxa. Carnivora by Gustafson (in progress); Ceratomorpha by Schiebout (in progress). G is Rancho Gaitan. TAXA K-Ar dates: 40.0 MARSUPALIA ?Peratherium sp. INSECTIVORA Geolabis n. sp. Lepictis douglasiLeptictis wilsoni Apternodus cf. brevirostris PRIMATES Omomyid gen. et sp. indet. Rooneyia viejaensis RODENTIA Leptotomus leptodus L. gigansMytonomys gaitaniaMicroparamys perfossusIschyrotomus cf. /. petersoniManitshajoahniculiIschryomys blacki Titanotheriomys veterior Cylindrodon fontis PseudocylindrodonneglectusPseudocylindrodon cf. neglectus P. texanus P. cf. texanus Ardynomys occidentalis Jaywilsonomysojinagaensis J. pintoensis J. aff. pintoensisAdjidaumo cf. A. minutus Viejadjidaumo magniscopulicf. Viejadjidaumo sp. indet. Aulolithomysbounites A. cf. A. bounites FAUNAS 38.6 36.8 31.4 Candelaria l.f. Porvenir l.f. Little Egypt l.f. Airstrip l.f. Ash Spring l.f. X X X X X X X X X G X X XX X X XX X XX X XX X G G X X X X X X [Table 14 continued, next page] 27 [Table 14, continued] TAX A FAUNAS K-Ar dates: 40.0 38.6 36.8 31.4 Candelaria l.f. Porvenir 1.f. Little Egypt l.f. Airstrip l.f. Ash Spring l.f. Meliakrouniomys wilsoni X Yoderimyslustrorum X Eutypomys inexpectatus X X cf.eutypomyidgen.etsp.indet X cf. Simimys sp. indet. X Subsumus candelariae X Prolapsus sp. X CARNIVORA Harpagolestes sp. X Ischnognathussavagei X Hemipsalodon n. sp. X Hyaenodon sp. X X X Miacis n. sp. X X Daphoenus sp. X Daphaenocyon sp. X Hesperocyon sp. X Fetid gen. et sp. indet. PER ISSODACTYLA Epihippus cf. E. gracilis X Haplohippus texanus X Mesohippustexanus X Mesohippussp. X PSthenodectesaustralis X Menodus bakeri X X Teleodussp. X Dilophodon n. sp. X Colodon cf. hancocki X Colodon n. sp. X X Hyracodon sp. X X X Amynodon sp. X Megamynodon sp. X Caenopus sp. X Schizotheroides jackwilsoni X X Toxotherium cf. T. woodi X ARTIODACTYLA Brachyhops wyomingensis X Archaeotherium cf. A. mortoni X X X Protoreodon petersoni X G P. pumiius X X Agriochoerus antiques X X A. maximus G Bathygenys reevesi XG Aclistomyctermiddletoni X [Table 14 continued, next page] 28 [ Table 14, continued] TAX A FAUNAS K-Ar dates: 40.0 38.6 36.8 31.4 Candelaria l.f. Porvenir l.f. Little Egypt l.f. Airstrip l.f. Ash Spring l.f. X Merycoidodontidaegen. et sp. indet. X G Merycoidodon dunagani X Limnenetes cf. L. platyceps X PProdesmatochoerus meekae X Leptoreodonmarshi X Poabromyluskayi X P. minor X Toromeryx marginatus X Heteromeryx dispar X Oromeryx sp. X Oromerycinae gen. et sp. indet. X Eotylopus cf. E. reedi Hidrosotherium transpecoensis X Poebrotherium franki X X Poebrotherium sp. X Hypertragulusheikeni G Leptomeryx defordi X Hypisodus cf. H. minimus X X 35.2 and 33.3 m.y. Correlations with this span are perhaps as close as can be hoped for with our present tools. Last appearances within the Porvenir l.f. are important. Schizotheroides is only known elsewhere from the Sage Creek Eocene of Montana. Protoreodon first appears in Uinta B in Utah and survives until the early Chadronian Porvenir l.f. Although the specimen is very fragmentary, Oromeryx is present in the Porvenir l.f. and is another survivor from Uinta B. The only presently known endemic forms seem to be Rooneyia, Haplohippus, Aclistomycter, and Hidrosotherium. Ischnognathus, in my opinion, is indeterminate. All other genera have been identified elsewhere. The succession of appearance of genera within the family Merycoidodontidae in the Vieja area is interesting. Merycoidodonts in the Porvenir l.f. are known only from a few indeterminate fragments from the Big Cliff area. In the southern area near Adobe Spring, remains of Aclistomycter are found in an interval 25-40 feet above the Buckshot Ignim brite but still beneath the lower marker bed. This is the lowest occurrence of an identifiable merycoidodont. In the overlying Little Egypt l.f. in the upper part of the Chambers Tuff about 60 feet below the Bracks Rhyolite, Merydoidon dunagani and Bathygenysreevesi are common. Limnenetes and PProdesmatochoerus appear in the Airstrip l.f. 570 feet above the Bracks Rhyolite in the Capote Mountain Tuff. The following are members of the Flagstaff Rim fauna as listed by Emry (1973) that are absent from the Porvenir l.f. A microfauna cannot be obtained by washingfrom the Chambers because of sediment consolidation, but a surprising number of insectivores, rodents, and the primate are known from skulls or jaw fragments. Peratherium is absent but I suspect this is an accident of collecting. The apatemyid Sinclairella and the Orders Chiroptera and Pholidata are absent. The family Castoridae and the order Lagomorpha also are absent. The carnivores, tapirs, and rhinos are still being studied so I do not wish to comment 29 beyond listing them in table 14. The families Leptochoeridae and Anthracotheriidae are not present in the Texas fauna. It is difficult to assess the significance of the absence of certain taxa in faunal lists. Their absence may be for a number of reasons: geographic distribution, temporal distribution, the accessibility of microfaunas, and length of time of collecting. Nonetheless, certain groups are conspicuous by their absence from the Porvenir l.f. No lagomorphhas been found in the Porvenir l.f. nor has anylagomorph been found in any west Texas fauna prior to the Arikareean. During 20 yearsof collecting it would be reasonable for one to assume that lagomorphs would have been discovered if they were present. Similarly, no pholidotan, leptochoerid, nor anthracothere has been found in west Texas. Like the lagomorphs, I would have thought they would have been found by now if they had been present. Although this is based on negative evidence I believe they were not present be cause of geographic distribution. Taxa that are present in the Porvenir l.f. but absent at Flagstaff Rim according to the preliminary list of Emry (1973) are the tar si iform Rooneyia, Leptotomus, Micropara mys, Ischryotomus, Manitsha, Pseudocylin drodon, Ardynomys, Viejadjidaumo, Au/o lithomys, Yoderimys, Eutypomys, cf. Simi mys, Haplohippus, Schizotheroides, Protoreo don Heteromeryx, Aclistomycter, and Hidro f sotherium. Of these, Leptotomus, Microparamys, and Manitsha are also present in the late Eocene as are Schizotheroides and Protoreodon. The latter five genera may represent a difference in temporal distribution and signify a slightly earlier age for the Porvenir or perhaps represent a geographic difference. I prefer the former because of the slight difference in potassium-argon dates. More important are the genera held in common, the most significant of which I believe to be Meliakrouniomys, Hemipsaiodon, Toxotherium, Brachyhyops, Eotylopus, and a small oromerycid. On the basis of these and other genera held in common, the lower part of the Flagstaff Rim and the Porvenir faunas are both early Chadronian. Table 15. Distribution of K-Ar dates in the Vieja area and northern Davis Mountains. 1. 40.0 ± 2.0, boulder in Gill Breccia. Gives lower limit for Colmena Tuff and Candelaria l.f. (Wilson et al. 1968). 2. 38.6 ± 1.2,BuckshotIgnimbrite. Givesupperlimitfor Colmena Tuff and Candelaria l.f. and lower limit for Chambers Tuff and Porvenir l.f. (Wilson et al. 1968). 3. 37.2 ± 0.7 Star Mt. Rhyolite and 36.6 0.7 Gomez Ash-Flow sheet (ave of 6 dates) ± ± 37.1 0.4 Adobe Canyon Fm. (ave of 2 dates) 36.2 ± 0.4 Sleeping Lion Fm. (ave of 2 dates) 35.6 ± 0.3 Barrel Springs Fm. (ave of 2 dates) ± 36.8 0.7 Goat Canyon Trachyte all give upper date for Huelster Fm. with Hyracodon (all in Parker and McDowell, in press). 4. 36.5 ± 1.2 and 36.8 Bracks Rhyolite. Gives upper date for Porvenir l.f.(Wilson et al. 1968). 5. 31.4 ± 0.5 Mitchell MesaRhyolite Tuff (=Brite Ignimbrite) average of five dates (pers. com. F. W. McDowell Sept. 21, 1976). This is the most widespread igneous unit in Trans-Pecos Texas. It is an upper date for the Airstrip l.f. in the Vieja area. 6. 22.3and 19.2averageagesonolderandyoungerdike swarm in the Vieja area. Dasch et al. 1969. Little Egypt Local Fauna —Most of the material that makes up this local fauna comes from two general localities. The most prolific. Reeves Bonebed, occurs approximately 30 feet below the upper marker bed in the Chambers Tuff. The other area is known as Chalk Gap Draw. The fauna contains the highest occurrence of Manitsha in the Vieja area. The other rodents indicate a correlation with Pipestone Springs, Montana. The presence of Bathygenys in both faunas supports this conclusion. Limnenetes is associated with Bathygenys, however, in the overlying Airstrip l.f. and although the faunas are separated by almost 600 feet of stratigraphic section, the time interval is apparently short. The presenceof the small species of Hyaenodon similar to those from Pipestone Springs in the Airstrip l.f. supports the same conclusion. Airstrip Local Fauna This fauna marksthe first occurrence of Poebrotherium, Limnenetes, PProdesmatochoerus, and Hypisodusin the Vieja area. Limnenetes is the commonform, in this local fauna followed by Poebrotherium, Bathygenys, and Hyaenodon. 30 Rancho Gaitan Local Fauna -Wood (1974) states that "the three rodents of the Rancho Gaitan local fauna do not fit in with any of the other Vieja local faunas" and suggests that the Rancho Gaitan l.f. falls on the Eocene-Oligocene boundary. The presence of Bathygenys and a medium-size oreodont, however, led Ferrusquia-V to correlate it with the Little Egypt l.f. Also, the association of Bathygenysand Agriochoerus maximus is known in the Pipestone Springs fauna of Montana and supports a somewhat later correlation. Hyper tragulus heikeni, according to Ferrusquia-V., is an intermediate between Simimeryx hudsoni from the Pearson Ranch l.f. of California and Hypertragulus calcaratus and is slightly more advanced than the former. Ferrusquia- V. (1967) and Wood (1974) point out the intermediate position of Jaywilsonomys to Pareumys from the Hartman Ranch l.f. and Sespemys of the late Oligocene. In short, I support Ferrusquia-V. in correlating the Rancho Gaitan l.f. as a whole with the Little Egypt l.f. and both in turn with Pipestone Springs, Montana. Paleoenvironment The Vieja Group is a succession of voicaniclastic sediments intercalated between flow rocks and ignimbrites. The Gill Breccia is a flow breccia, perhaps consisting of several flows. Its source lay to the south or southwest according to Walton (1972) and was probably not far away. In places a smooth erosion surface can be seen at the top of the Gill. The lower or sedimentary facies of the Colmena consists of sorted sandstone, described by Walton (1972) as a volcanic litharenite. Cross-bedding is found in the lower facies and suggests braided stream deposits. Gastropod steinkerns are common and a few plant impressions have been found in this facies. Fragments of crocodile or alligator are present but not common. The area near the mouth of Capote Creek during the time of accumulation of the lower facies was a broad valley occupied by braided streams that flowed between active volcanoes. Volcanic activity was closer during the time of the ac- Table 16.Numbersofspecimensofmediumsizedherbivores, Porvenir l.f. FMNH TMM OU TOTAL MesohippusAgriochoerusLep tomeryxHeteromeryxProtoreodon 36 31 5+ 7 3 37 26 27 5 8 0 6 6 0 0 81 63 38+ 12 11 PoabromylusAclistomycterMerycoidodontidae HaplohippusBrachyhyopsArchaecotherium 7 0 1 2 1 1 5 6 1 0 0 0 2 0 0 0 0 0 7 6 2 2 1 1 Schizotheroides 0 1 0 1 Oromeryx 0 1 0 1 cumulation of the lahars and the upper glassfacies. The latter is the result of nearby explosive volcanic activity. Shards and pumiceform most of the matrix. Sedimentary structures are very rare. Only one concentration of bones, Clabaugh's Bonebed, was found in this facies. The environment was probably one of the intermittent accumulation of volcanic ash. At times the ash falls may have temporarilykilled off the lower vegetation, forcing the retreat of the animals dependent on it, but as the tuffaceous soil weathered the plantsquickly reestablished and the animals returned. The environment in the Vieja area duringthe time of accumulation of the Chambers Tuff was more tranquil. Coarse conglomeratescontain cobble-sized material of reworked Cretaceous limestone that came from the Laramide folded uplands to the west in Mexico. These channel sandstones and conglomerates often contain bone and tooth fragments. At a locality known as "red mound" one of these channels grades laterally into a bright red mudstone that probably representsoverbank deposits. These deposits contain coprolite-shaped masses with bone and enamel chips, and what appear to be burrows. Most of the material from the red mound locality consisted of small forms (table 5) but identifiable fragments of larger forms were also found. Elsewhere mudstones are chocolate brown and light brown to reddish 31 brown and show very little bedding. A particular mudstone unit may be from a few inches up to 20 feet (16 m) thick. These show very little bedding except at color changes. Individual bones and teeth, some rodent-gnawed, are widely scattered in a dense, very hard mudstone but no articulated specimens were found. The mudstone probably represents slowly accumulating volcanic ash rich soils. No alligator or crocodile remains have been found. Turtles are present but not common in the Chambers Formation, but dermal scutes of lizards are fairly common. Gastropods identified as Helix leidyi are locally common in both the Colmena and Chambers. In the lower part of the Chambers, gastropod steinkerns three inches (7.6 m) in diameter have been found. At one locality near Rancho Gaitan in Mexico gastropods occur by the thousands. They are one to two inches at their widest diameter. At one locality in the Chambers approximately 10 feet (3 m) of very hard light greentuff is exposed. The tuff was deposited as a mud flat near the western limit of the Bracks Rhyolite. It may be in either the top of the Chambers or the base of the Capote Mountain. Trackways are found at several levels in the tuff. Because of the extreme hardness and lack of good bedding it was beyond our means to remove them, so flexible molds were made. One large mold about 9 by 18 feet (2.7—5.5 m) contained trackways of a variety of forms. One was of a bird about the size of a pigeon and another about the size of a turkey; neither is represented by skeletal material. The largest footprints must have been those of a titanothere. Two trackways were made by carnivores, one large that could belong to Hyaenodon and the other small that could belong to either Daphoenus or a miacid. A trackway of medium-size clawed form might belong to Agriochoerus. A trail of small artiodactyl footprints is the right size for Bathygenys. Medium-size footprints with three distinct toes are either tapir or rhino. All of these are represented on the largestmold. Smaller molds contain the trail of an arthropod the size of a large scorpion, a small turtle, and a small three-toed trackway that probably was left by Mesohippus. Such a concentration of animals in one small area must mean that it was close to a watering area. No impressions of leaves are seen, nor was there any petrified wood nearby. It may be that the area was part of a broad mud flat adjacent to a river or intermittent stream. Raindrop impressions and filled mudcracks are present on the bedding surfaces. The paleoenvironment of the Chambers Tuff seems to have been more open than that of the Colmena, more removed from direct volcanic activity and probably dryer. A small fossil flora was described by Berry(1919) from the basal tuffs of the Huelster Formation in the Barilla Mountains, a northeastern extension of the northern Davis Mountains. The Hyracodon tooth reported on pages 1, 24 also came from the lower tuffs of the Huelster Formation. The Huelster is correlated with the Chambers Tuff on the basis of the K-Ar dates of the Bracks Rhyolite of the Vieja area with similar dates on the Star Mountain Rhyolite (table 16) of the McCutcheon Group (Eifler 1951). The fossil flora from the Barilla Mountains has been reinterpreted by Axelrod and Bailey(1976, p. 237): “The Barilla flora has the fan-palm (Saba- Htes), feather-palm (Geonomites), and abundant laurel (Ocotea, recorded as Oreodaphne), evergreen holly (Hex) and 2 or 3 others that were not illustrated, including a reported walnut (Juglans). To judge from the small sample, it implies a warmer climate than the Bernalillo (New Mexico) flora which is consistent with its position 560 km southeast and probably lower (300m) altitude. Representing the upper part of a dry mixed sub tropical flora it finds analogy with mixed vegetation in southern Tamaulipas and southward along the east front of the Sierra Madre Oriental." Differences in elevation between the present day position of the Huelster and Chambers are the result of mid-to late-Tertiaryfaulting but during the early Oligocene the 32 two areas were probably close to the same elevation. In both areas the early Tertiaryrocks lie conformably on late Cretaceous. In the Barrillas, the McCutcheon Group lies on marine Cretaceous correlated with the Taylor marl of central Texas; in the Vieja area the Vieja Group lies on rocks as young as non-marine Cretaceous beds correctable with the Aguja and Javelina Formations of BigBend National Park. Axelrod and Bailey'sanalysis of the climate for the Barrilla flora also can be applied to the climatic conditions in the nearby Vieja area. Harris (1967a) interpreted a calichification of the tuffaceous fossiliferous sediment at Ash Spring to mean subarid conditions. He says “No typical caliche structures were seen in the thin sections but the preponderance of sugary fine-grained calcite and the friable nature of the rock are suggestive of calichification (personal communication, R. L. Folk, October, 1966). That the caliche was implaced in periods of subareal exposureduring the accumulation of the rock, rather than subsequently, is indicated by thin caliche beds in the upper part of the unit. .., and by .“ .. pebbles of caliche in a conglomerate . The Ash Spring l.f. is apparently the youngest of the Vieja Group and Harris (1967a) interpreted a progressive drying of the climate in the Vieja area. The Airstrip l.f. lived on the periphery of the volcanic area where tuff was accumulating. The deposits that contain the fossils show very few sedimentary structures. Some faint cross-bedding can be found and some of the beds fine upward so that a water environment for deposition seems certain. Turtles identified as Sty/emys sp. are common in the Capote Mountain whereas turtles are rare in the Chambers. Gastropods have not yet been found in association with vertebrates in the Capote Mountain. The Capote Mountain and the deposits at Ash Spring seem to have been laid down under drier conditions than those in the Colmena, and the Chambers representsconditions somewhat in between. This change was gradual over a period of six to eight m.y. Chronostratigraphy and the Chadronian-Uintan Boundary Throughout this report I have used the terms Uintan and Chadronian, avoiding use of the term Duchesnean. All three terms were defined by Wood et al. (1941) and have been in use ever since. There has been criticism of the way in which Wood et al. (1941) set up their units, and various authors have interpret ed them in different ways. For example, the Uintan age was defined as follows: “Uintan age—new provincial time term, based on the Uinta formation of northern Utah, i.e., the time of deposition of Uinta A-C, inclusive, with the included faunas." Some authors have interpreted this strictly to mean that the Uintan age is coincidental with the geochronof the Uinta Formation. Other authors have interpreted it to be the time during which the animals lived that characterize the Uinta Formation. The two do not necessarily have equivalent time spans. I cannot believe that Wood et al. (1941) meant to have the first interpretation strictly applied and indeed they so state (p. 6): "... the ages are not necessarily coextensive with their types, and the precise limits between successive ages are intended to be somewhat flexible and may presumably be modified in the light of later discoveries. Thus the Wasatchian age is more extensive that the known mammalian faunas of the type Wasatch and probably less extensive than the time equivalent of the entire Wasatch groupin the type area. The Barstovian age includes units which may be older than any part of the Barstow formation, and are certainly older than any of the faunules now known from Barstow." Wood et al. (1941) define Chadronian as "Chadronian age new provincial time term, - based on the Chadron formation, type locality near Chadron, Nebraska; type area, northwestern Nebraska and southwestern South Dakota; includes the old term "Titanotherium beds," used in its most extended sense. It may also be defined, faunally, as the time 33 during which Mesohippus and titanotheres coexisted." The basis for the Chadronian age is by this definition four-fold: (1) a geochronof the type locality (but see Schultz and Stout 1955, and Harksen and Macdonald 1969 for two additional "type localities;" (2) time of accumulation of deposits in a type area; (3) the time of accumulation of beds containing "Titanotherium;" and (4) a biologic definition based on time of coexistence of Mesohippus and titanotheres. Such a definition does not support the interpretationthat the Chadronian age "is based" on the geochron of the Chadron Formation. Wood et al. (1941) also established the Duchesnean age. It was "based on the Duchesne River formation of northeast Utah." This was an attempt to fill a gap in the paleontological record that had long been recognized (Matthew 1924, p. 750; Osborn 1929, p. 99; Simpson 1933, p. 84). Wood et al. (1941) go on to say that "A faunal definition would be premature except to note the abundance of Te/eodus. Hence the faunas are listed separately." They then list the Duchesne River fauna and a correlative fauna, California Institute of Technology Locality 150 in the Sespe of California. In the central Rocky Mountain area two sections were thought to have sediments depositedduring a time interval between Uintan and Chadronian. These were at Beaver Divide, Wyoming and the Duchesne River Formation, Utah. The problems connected to the former area, only recently, have been cleared up in Emry (1975). The vertebrate fauna from the Duchesne River Formation was collected by 0. A. Peterson, J. L. Kay, and John Clark of the Carnegie Museum and described during the 19305. Scott (1945) brought together the various descriptions of the fauna, and added Came/odon from the Beaver Divide Uintan, to constitute a faunal basis for the Duchesnean age. The stratigraphy of the Duchesne River Formation has recently been studied by Andersen and Picard (1972) who also give the distribution of the vertebrate fossils within the formation. The faunal lists given by Andersen and Picard (1972) with the addition of Brachyhyops (Wilson 1971b) are the most recent faunal characterization of the Duchesne River Formation. My reluctance to use the term Duchesnean was caused by the uncertainty of what constituted the faunal basis for Duchesnean age. In order to be useful for correlation beyondthe limits of the type area the Duchesnean should have a characteristic fauna. As Ted- ford (1970, p. 692) succinctly states, ". re .. vision of mammal 'Ages' depends on biological evidences, as these temporal units cannot serve if restricted to the geochron of litho ... stratigraphic units." The Duchesnean as originally used by Wood et al. (1941) and Scott (1945) included the faunas of the Randlett, Halfway, and Lapoint members of the Duchesne River Formation. Since that time, as will be shown below, the Randlett and the Halfway have been placed in the Uintan, on faunal evidence, leaving only the Lapointfauna to characterize Duchesnean. Subsequently, Clark et ai. (1967) divided the Duchesnean into an early "La Pointian" and a later Viejan. Their Viejan is characterized bythe Porvenir l.f. which I, for reasons stated below, assign to the Chadronian, so once again, the Lapoint fauna is the only one left to characterize Duchesnean. A brief resume of the history since Scott (1945) of pertinent faunal elements used to characterize the Duchesnean age follows. Kay (1934) divided the Duchesne River Formation of Utah into three "horizons" which he named Randlett, Halfway, and La- point from bottom to top. More recentlyAndersen and Picard (1972) divided the Duchesne River Formation into four Members, Brennan Basin, Dry Gulch Creek, Lapoint, and Starr Flat. The Brennan Basin Member contains the fauna attributed to the Randlett; the Dry Gulch Creek contains the fauna attributed to the Halfway, and the LapointMember contains the fauna attributed to the Lapoing horizon. The Starr Flat Member is non-fossiliferous. As faunas of either the late Eocene or early Oligocene in the western interior of North America and California became better known the tendency has been to divide the Duchesnean. Gazin as early as 1955 (chart 1) 34 recognized that the fauna from the Randlett or Brennan Basin Member was so closely related to the nearby late Eocene Myton fauna that he included the Randlett within the late Eocene Uintan Age. Gazin (1956; 1959) reaffirmed this opinion, which was based primarily on his study of the late Eocene artiodactyls. A similar position was reached by Radinsky (1963) from his study of the tapiroids. In regard to the tapiroid Dilophodon leotanus from the Randlett, he says: "Comparison between Dilophodon specimens from Uinta C, Randlett, Badwater and Sage Creek faunas suggest that they are all conspecific." Clark et al. (1967), Wilson et al. (1968), Tedford (1970), and Golz (1976) all followed Gazin (1955) and included the Randlett fauna with Uintan. The faunas of the Halfway or Dry Gulch Member and the Lapoint Member would then be left to characterize Duchesnean time. The fauna from the Dry Gulch, according to Andersen and Picard (1972), consists of two taxa. Eosictis avinoffi, known from a singlefragmentary C-P4, and was described by Scott (1945) as a felid. A single specimen as fragmentary as the type of Eosictis is not particularly useful for purposes of correlation. Epihippus intermedius is the other taxon that represents the Dry Gulch fauna as listed byAndersen and Picard (1972). Because all other species of Epihippus occur in the Uintan I would prefer to include the Halfway fauna as Uintan as did Simpson (1933; 1946). Other authors list additional taxa from the Halfway. Peterson (1934, p. 383, fide Kay) saysthatMesagriochoerusprimus(CM 11904) "was taken from the upper part of the Halfway Horizon." Mesagriochoerus was synonymized with Protoreodon pumilus by Wilson (1971a). P. pumilus is found in both Uintan and Chadronian (Wilson 1971a). Scott (1945) lists ?Protitanotherium (CM 11, 996) as having come from the Halfway and says, "It is entirely possible that this fragment should be referable to one of the characteristic Uinta genera, Protitanotherium Hatcher or Diplacodon Marsh. Between these two, it is not feasible to make a definite choice because the fragment might equally well belong to either genus ..." So the only useful taxon in the Halfway faunas at present for correlation purposes is Epihippus and its affinities are with the Uintan. For this reason I would follow Clark et al. (1967) and McKenna et al. (1973) and include both the Brennan Basin (Randlett) and Dry Gulch (Halfway) within the Uin tan. This left only the Lapoint fauna to characterize a Duchesnean time span. Clark et al. (1967, p. 56-59) in their monograph on the Chadron Formation of South Dakota discuss their faunas in relationship to pre-Chadronianfaunas elsewhere including "the Vieja fauna." The Vieja fauna as used by Clark et al. (1967) is now limited to the Porvenir I. f. of this report. Clark et al. (1967, p. 59) summarize their conclusions as follows. I have inserted in brackets the term Porvenir l.f. to emphasizethat I regard the term "Vieja fauna" as being too inclusive. "The Chadron includes three distinct rock- time units with characteristic faunal associations (see Fig. 25). The oldest, the Ahearn, is apparently younger than the Vieja [Porvenir] which may be regarded as late Duches nean .... "The lower two members of the Duchesne River Formation, however, are probably Eocene and include the typical Eocene speciesof Protoreodon and Diplobunops but are somewhat younger than Uinta C. the upperDuchesne River, the La Point, includes the titanothere, Teieodus, which is associated with Mesohippus viejensis [M. texanus McGrew (1971)] and Agricochoerus [sic] in the Vieja [Porvenir l.f.] and with Mesohippusin the Cypress Hills. The rest of the known La Point fauna is either transitional between the Uintan and Chadron forms or is represented by specimens so inadequate that their relationships cannot be precisely determined. "The La Point Member of the Duchesne River Formation may then be properly considered typical of Duchesnean time (type lo - cality, 12 miles west of Vernal, Utah Kay, 1934) with the understanding that no rock section described up to the present repre 35 sents all of Duchesnean time, that the LaPoint fauna is early Duchesnean, and that the ViejaFormation of Texas (Stoval [sic], 1948) is late Duchesnean and should be considered the type for this part of Duchesnean time. Teleodus, associated with Poabromylus, Epihippusuintensis, and Diplobunops, indicates La Pointian or early Duchesnean; and the Teleodus- Agriochoerus-Mesohippus viejensis [seeabove] association indicates Viejan [Porvenir] or late Duchesnean age." Several points in the quotation above need further comment. First, Stovall (1948) does not describe a Vieja Formation, nor has anyone else. Second, I disagree with their creating new "ages" based on the geochrons of lithologic units. Mesohippus viejensis as emphasized by McGrew (1941), is tied to South Dakota specimens and, as Tedford points out (1970), Clark et al. (1967) "are incorrect in characterizing their 'Lapointian' with Epihippus and Diplobunops, neither of which is recorded from Lapoint." Clark's assignment of the Porvenir l.f. to the Duchesnean was premature because the Porvenir l.f. presently contains three generaof insectivores, five genera each of rodents and carnivores, thirteen genera of perissodactyls, and six of artiodactyis, a total of thirty- two genera that occur in common with Chadronian faunas to the north. For this reason I have assigned the Porvenir l.f. to the Chadronian. Tedford's (1970, fig. 6) chart shows the occurrence of mammal species recorded from the Duchesne River Formation. More recent information of the occurrence of some of the genera, particularly those species from the La- point, modified his chart. Most authors would correlate the Myton and the Randlett so that they would not appear to be as widely separated as shown by Tedford. Pentacemylus is now known to be present in the Chadron Formation, as is Poabromylus kayi. Brachyhyopsis present in the Lapoint, Flagstaff Rim, and the Vieja, as well as Beaver Divide. Hesso/estes, as will be reported later, extends down to the Uintan in the Agua Fria area (fide Gustafson). Dyscritochoerus is so fragmentary that, in my opinion, it is not identifiable. The Eocene artiodactyis of southern California have recently been studied by Golz (1976). He gives in his table 4 the North American artiodactyl assemblages of Duchesnean Age and lists the genus Simimeryx represented by S. hudsoni in California and S. minutus in the Lapoint, Utah. The closely re lated form Hypertragulus heikeni was described by Ferrusquia-V. (1969) from the early Oligocene of Chihuahua Mexico. It is an intermediate between S. hudsoni and H. caicaratus, and very close to S. hudsoni and S. minutus and smaller than either. H. heikeni is associated with Protoreodon peterson, Agriochoerus maximus, cf. Leptomeryx, and Bathygenys alpha. The last three occur in Chadronian faunas in Montana and Wyoming. Small species of hypertragulids extend from the Lapoint into the Chadronian. Protoreodon pumilus is also listed by Golz (1976) in the artiodactyl assemblages of Uinta B and early "Uinta C" and "Randlett" as well as in the Duchesnean. In addition, I (Wilson 1971b) have found P. pumilus in the Chadronian so this taxon is not a useful time marker. Poabromylus kayi from the Lapoint fauna was compared by Wilson (1974) withPseudoprotoceras longinaris from the Chadronian and the two were synonymized. Golz (1976, table 4) also lists Protylopus pearsonensisfrom the Pearson Ranch l.f. of California as characteristic of the Duchesnean but the species has not been found elsewhere. The North American artiodactyl assemblage of Duchesnean age as used by Golz is not very helpful for correlation but it must be remembered that he, following Gazin (1956), regarded the Brennan Basin or Randlett as indistinguishable from the Uintan, and his "North American artiodactyl assemblages of Duchesnean age" represent onlyfaunas considered by him to correlate with the Lapoint (Golz 1976, fig. 4). The Porvenir l.f. has only four genera, Microparamys, Hyaenodon, Teleodus, and Protoreodon in common with the Pearson Ranch l.f. of California, which was placed in the Duchesnean by Wood etal. (1941). Recent work 36 by Coombs (1971) shows that all specimensof Simidectes from Utah are Uintan and its range to Duchesnean is based on its presence at Locality 150 of the Pearson Ranch l.f. Pterodon californicus also from the Pearson Ranch l.f. was made a junior synonym of Hyaenodon vetus by Mellett (1977). Unfortunately, the specimen of Hyaenodon from the Lapoint is only fragmentary RMT2 ancj could only be referred to Hyaenodon, cf. H. vetus by Mellett (1977, p. 20). Another specimen that Mellett says is of uncertain allocation but was tentatively referred to Hyaenodon, cf. H. vetus, is from the Chadronian of Flagstaff Rim, Wyoming. Wilson and Szalay (1976) report the association of Hyaenodon with Mahgarita stevensi, Protoreodon pumilus, and Leptoreodon in the “Skyline conglomerate" of the Agua Fria area. Gustafson (in progress) has identified this Hyaenodon as H. cf. vetus. With this revision, of the 12 taxa listed byTedford (1970, fig. 6) from the Lapoint, only Simimeryx minutus is unique, five continue from the Randlett or earlier, and eight appear in the Lapoint and are also found in the Chadron Formation. Tedford, however, (1970, p. 692) goes on to suggest that: "Teleodus, together with the first appearance in the Lapoint of Hyaenodon, Hyracodon, Poabromylus and Simimeryx, might constitute the faunal basis for a mammalian age distinct from the Uintan Both Scott (1945) and Radinsky .. . (1967) question the identification of the specimen of Hyracodon primus and also the specimen described as Mesamynodon medius, both from the Lapoint; so that the first certain appearance of Hyracodon is Chadronian. In addition, it would be valuable to have some statistical basis to distinguish Teleodus from Menodus. At present, only Hyaenodon, Poabromylus, and Simimeryx have their first appearance in the Lapoint and are useful as the faunal basis for a Duchesnean Mammal Age. If the Candelaria and Porvenir l.f. are correctly assigned to the Uintan and Chadronian respectively, a Duchesnean fauna is absent in the Vieja area. The Candelaria l.f. and the Porvenir l.f. have only three genera in common, Ischyrotomus, Manitsha, and Protoreodon, whereas the Porvenir l.f. has six genera, Pareumys, Hyaenodon, Teleodus, Brachyhyops, Protoreodon, and Poabromylus in common with the Lapoint A Lapointian fauna should therefore be intermediate. If so, my previous correlation of the Porvenir and Lapoint would be in error. However, the six genera listed above that occur in both the Lapoint and Porvenir faunas constitute all but two of the adequately represented mammalian taxa of the Lapoint fauna. Murphy (1977) and Woodburne (1977) urge that one taxon be used to identify the lower boundary of a chronostratigraphic unit. Such a practice was suggested by Simpson(1933). Additional taxa help to confirm the correlation but as pointed out strongly byWoodburne (1977) and Murphy (1977), the use of more than one taxon to identify a chronostratigraphic unit results in diminished precision. I therefore urge that the first appearance of Mesohippus be used to mark the beginning of Chadronian time. FollowingMurphy (1977) and Woodburne (1977) I do not propose an upper limit because it would depend on the first appearance of another taxon for the overlying chronostratigraphicunit and such is beyond the scope of this report. I urge Mesohippus for a number of reasons. It is widespread; it is common, and throughthe work of recent authors (Clark et al. 1967; McGrew 1971; Forsten 1970, 1971a, 1971b) is well known. In contrast the titanotheres have not been restudied for almost 50 years. If this practice is followed, the Porvenir l.f. is Chadronian and the first appearance of the primitive species Mesohippus texanus McGrew (not M. viejensis Clark and Beerbower) marks the beginning of Chadronian time. 37 SUMMARY AND CONCLUSIONS The stratigraphic positions of the Candelaria, Porvenir, Little Egypt, and Airstriplocal faunas are given within the Vieja Group. It can be demonstrated in the field that, although the area is complexly faulted, these four local faunas occur in supt.position. The Candelaria l.f. is late Uintan and occurs primarily in a lower sorted facies of the Colmena Tuff. A K-Ar date of 40.0 ± 5.0 was obtained for a volcanic rock in the underlying Gill Breccia, and 38.6 is believed to be the best date for the overlying Buckshot Ignimbrite. The Porvenir l.f. (Viejan fauna of others) occurs in the lower part of the Chambers beneath an upper marker bed and above the Buckshot Ignimbrite. Thirty-two genera in the Porvenir l.f. occur in common with Chadronian faunas to the north and six are in common with the Lapoint fauna. The Little Egypt l.f. is Chadronian and is overlain bythe Bracks Rhyolite dated at approximately36 m.y. The Airstrip l.f. is found in the up per part of the Capote Mountain Tuff, which in turn is overlain by the Mitchell Mesa Rhyolite (= Brite Ignimbrite) of approximately31 m.y. The latter unit is widespread over the west Texas volcanic field. Other local faunas, the Ash Spring, an unnamed one in the Garren Group, and the Rancho Gaitan l.f. of Chihuahua Mexico are not in superposition but are correlated to the other local faunas on biologic criteria. The only basis for long distance correlation in the absence of K-Ar dates is biological. Unfortunately, the Duchesnean is poorlycharacterized biologically so that close correlation is difficult. The dates of 39.3 ± 0.8 on a tuff at the contact of the Halfway and Lapoint in Utah and the date of 38.6 ± 1.2 for the Buckshot Ignimbrite in Texas are close. Hopefully, a sufficiently distinctive fauna to warrant a mammalian “age" called Duchesnean can be found. I urge that the base of the Chadronian be marked by the first appearance of Mesohippus. 38 REFERENCES Andersen, D. W., and M. D. Picard, 1972. Stratigraphy of the Duchesne River Formation (Eocene-Oligocene?), northern Uinta Basin, northeastern Utah. 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